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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Cladus: Sarcopterygii
Cladus: Rhipidistia
Cladus: Tetrapodomorpha
Cladus: Eotetrapodiformes
Cladus: Elpistostegalia
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Classis: Reptilia
Cladus: Eureptilia
Cladus: Romeriida
Subclassis: Diapsida
Cladus: Sauria
Infraclassis: Lepidosauromorpha
Superordo: Lepidosauria
Ordo: Squamata
Cladus: Unidentata, Episquamata
Cladus: Toxicofera
Subordo: Serpentes
Infraordo: Caenophidia
Superfamilia: Colubroidea

Familia: Colubridae
Subfamilia: Colubrinae
Genus: Lycodon
Species (59): L. albofuscus – L. alcalai – L. anamallensis – L. aulicus – L. banksi – L. bibonius – L. butleri – L. capucinus – L. cardamomensis – L. carinatus – L. cathaya – L. cavernicolus – L. chrysoprateros – L. davidi – L. deccanensis – L. dumerili – L. effraenis – L. fasciatus – L. fausti – L. ferroni – L. flavicollis – L. flavomaculatus – L. flavozonatus – L. futsingensis – L. gammiei – L. gibsonae – L. gongshan – L. hypsirhinoides – L. jara – L. kundui – L. laoensis – L. liuchengchaoi – L. mackinnoni – L. meridionalis – L. muelleri – L. multifasciatus – L. multizonatus – L. namdongensis – L. ophiophagus – L. orientalis – L. paucifasciatus – L. pictus – L. rosozonatus – L. rufozonatus – L. ruhstrati – L. sealei – L. semicarinatus – L. septentrionalis – L. sidiki – L. solivagus – L. stormi – L. striatus – L. subcinctus – L. synaptor – L. tessellatus – L. tiwarii – L. travancoricus – L. zawi – L. zoosvictoriae

Lycodon Boie in Fitzinger, 1826: 29, 57

Type species: Coluber aulicus Linnaeus, 1758, by original designation.


Cercaspis Wagler, 1830: 191
Type species: Hurria carinata Kuhl, 1820, by monotypy.

Dinodon Duméril, Bibron & Duméril, 1854: 447
Type species: Dinodon cancellatum Duméril, Bibron & Duméril, 1854, by monotypy.

Euprotodon Cope, 1860: 262
Type species: Coluber aulicus Linnaeus, 1758, by original designation and monotypy.

Eumesodon Cope, 1860: 262
Type species: Eumesodon semicarinatus Cope, 1860, by original designation.

Tytleria Theobald, 1868: 66
Type species: Tytleria hypsirhinoides Theobald, 1868, by monotypy.

Primary references

Fitzinger, L.J. 1826. Neue classification der reptilien nach ihren natürlichen verwandtschaften : nebst einer verwandtschafts-tafel und einem verzeichnisse der reptilien-sammlung des K. K. zoologischen museum's zu Wien. J.G. Heubner: Wien. BHL
Wagler, J.G. 1830. Natürliches System der Amphibien, mit vorangehender Classification der Säugetiere und Vögel. Ein Beitrag zur vergleichenden Zoologie. J.G. Cotta'schen Buchhandlung: München, Stuttgart, & Tübingen. 354 pp. BHL Reference page.
Duméril, A.M.C., Bibron, G. & Duméril, A.H.A. 1854. Erpétologie générale ou histoire naturelle complète des reptiles. Tome septième. Première partie. Librarie encyclopédique de Roret, i-xv + 780 pp. BHL
Cope, E.D. 1860. Catalogue of the Colubridae in the Museum of the Academy of Natural Sciences of Philadelphia, with notes and descriptions of new species. Part 2. Proceedings of the Academy of Natural Sciences of Philadelphia 12: 241–266. BHL Reference page.
Theobald, W. 1868. Catalogue of Reptiles in the Museum of the Asiatic Society of Bengal. Journal of the Asiatic Society of Bengal. Part 2. Natural History 37(extra number 146): vi + 7–88. BHL

Additional references

Daltry, J.C. & Wüster, W. 2002. A new species of Wolf Snake (Serpentes: Colubridae: Lycodon) from the Cardamom Mountains, Southwestern Cambodia. Herpetologica 58(4): 498–504. DOI: 10.1655/0018-0831(2002)058[0498:ANSOWS]2.0.CO;2 Paywall Reference page.
Vogel, G. & Brachtel, N. 2008. Contribution to the knowledge of Lycodon ruhstrati (Fischer, 1886) in Vietnam - taxonomy and biology of a little-known species. Salamandra 44(4): 207–224. PDF Reference page.
Vogel, G., David, P., Pauwels, O.S.G., Sumontha, M., Norval, G., Hendrix, R., Vu, N.T. & Ziegler, T. 2009. A revision of Lycodon ruhstrati (Fischer 1886) auctorum (Squamata Colubridae), with the description of a new species from Thailand and a new subspecies from the Asian mainland. Tropical Zoology 22(2): 131–182. Reference page.
Vogel, G. & david, P. 2010. A new species of the genus Lycodon (Boie, 1826) from Yunnan Province, China (Serpentes: Colubridae). Bonn zoological Bulletin 57(2): 289–296. PDF Reference page.
Vogel, G. & Luo, J. 2011. A new species of the genus Lycodon (Boie, 1826) from the southwestern mountains of China (Squamata: Colubridae). Zootaxa 2807: 29–40. Preview. DOI: 10.11646/zootaxa.2807.1.2 Paywall Reference page.
Zhang, J., Jiang, K., Vogel, G. & Rao, D. 2011. A new species of the genus Lycodon (Squamata, Colubridae) from Sichuan Province, China. Zootaxa 2982: 59–68. Preview. DOI: 10.11646/zootaxa.2982.1.6 Paywall Reference page.
Vogel, G., Nguyen, T.Q. Kingsada, P. & Ziegler, T. 2012. A new species of the genus Lycodon Boie, 1826 from Laos (Squamata: Colubridae). North-Western Journal of Zoology 8(2): 344–352. PDF Reference page.
Vogel, G. & Harikrishnan, S. 2013. Revalidation of Lycodon hypsirhinoides (Theobald, 1868) from Andaman Islands (Reptilia: Serpentes: Colubridae) Taprobanica 5(1): 19–31. DOI: 10.4038/tapro.v5i1.5657 Open access Reference page.
Wostl, E., Hamidy, A., Kurniawan, N. & Smith, E.N. 2017. A new species of Wolf Snake of the genus Lycodon H. Boie in Fitzinger (Squamata: Colubridae) from the Aceh Province of northern Sumatra, Indonesia. Zootaxa 4276(4): 539–553. DOI: 10.11646/zootaxa.4276.4.6. Reference page.
Luu, V.Q., Bonkowski, M., Nguyen, T.Q., Le, M.D., Calame, T. & Ziegler, T. 2018. A new species of Lycodon Boie, 1826 (Serpentes: Colubridae) from central Laos. Revue suisse de Zoologie 125(2): 263–276. DOI: 10.5281/zenodo.1414221 Open access Reference page.
Vogel, G. & David, P. 2019. A new species of the Lycodon fasciatus complex from the Khorat Plateau, eastern Thailand (Reptiles, Squamata, Colubridae). Zootaxa 4577(3): 515–528. DOI: 10.11646/zootaxa.4577.3.6 Paywall Reference page.
Luu, V.Q., Ziegler, T., Ha, N.V., Le, M.D. & Hoang, T.T. 2019. A new species of Lycodon Boie, 1826 (Serpentes: Colubridae) from Thanh Hoa Province, Vietnam. Zootaxa 4586(2): 261–277. DOI: 10.11646/zootaxa.4586.2.3 Paywall Reference page.
Ganesh, S.R., Deuti, K., Punith, K.G., Achyuthan, N.S., Mallik, A.K., Adhikari, O. & Vogel, G. 2020. A new species of Lycodon (Serpentes: Colubridae) from the Deccan Plateau of India, with notes on the range of Lycodon travancoricus (Beddome, 1870) and a revised key to peninsular Indian forms. Amphibian & Reptile Conservation 14(3): 74–83. Reference page.
Wang, J., Qi, S., Lyu, Z.-T., Zeng, Z.-C. & Wang, Y.-Y. 2020. A new species of the genus Lycodon (Serpentes, Colubridae) from Guangxi, China. ZooKeys 954: 85–108. DOI: 10.3897/zookeys.954.53432 Open access Reference page.


Uetz, P. & Hallermann, J. 2023. Lycodon . The Reptile Database. Accessed on 22 June 2018.
Lycodon – Taxon details on Integrated Taxonomic Information System (ITIS).

Vernacular names
Deutsch: Wolfzahnnattern
English: Oriental Wolf Snakes
ไทย: งูปล้องฉนวน

Lycodon is a genus of colubrid snakes, commonly known as wolf snakes.[3] The Neo-Latin name Lycodon is derived from the Greek words λύκος (lykos) meaning wolf and οδόν (odon) meaning tooth,[4] and refers to the fang-like anterior maxillary and mandibular teeth.[2] They are nonvenomous, but many members of this genus strongly resemble the venomous kraits in appearance, an example of Batesian mimicry.

The genus Lycodon comprises 73 recognized species.[5]

Lycodon albofuscus (A.M.C. Duméril, Bibron & A.H.A. Duméril, 1854)
Lycodon alcalai Ota & Ross, 1994
Lycodon anakradaya Nguyen, Duong, Wood, Grismer, 2022
Lycodon anamallensis Günther, 1864
Lycodon aulicus (Linnaeus, 1758)
Lycodon banksi Luu et al., 2018
Lycodon bibonius Ota & Ross, 1994
Lycodon bicolor (Nikolsky, 1903) - Mackinnon's wolf snake
Lycodon butleri Boulenger, 1900
Lycodon capucinus F. Boie, 1827
Lycodon cardamomensis Daltry & Wüster, 2002
Lycodon carinatus (Kuhl, 1820)
Lycodon cathaya J. Wang, Qi, Lyu, Zeng & Y.-Y. Wang, 2020
Lycodon cavernicolus Grismer et al., 2011
Lycodon chapaensis (Angel & Bourret, 1933)
Lycodon chithrasekarai Wickramasinghe et al., 2020
Lycodon chrysoprateros Ota & Ross, 1994
Lycodon davidi G. Vogel et al., 2012
Lycodon davisonii (Blanford, 1878)
Lycodon deccanensis Ganesh, Deuti, Punith, Achyuthan, Mallik, Adhikari & G. Vogel, 2020
Lycodon dumerili (Boulenger, 1893)
Lycodon effraenis Cantor, 1847
Lycodon fasciatus (J. Anderson, 1879)
Lycodon fasciolatus (Shaw, 1802)
Lycodon fausti Gaulke, 2002
Lycodon ferroni Lanza, 1999
Lycodon flavicollis Mukherjee & Bhupathy, 2007
Lycodon flavomaculatus Wall, 1907
Lycodon flavozonatus (Pope, 1928)
Lycodon futsingensis (Pope, 1928)
Lycodon gammiei (Blanford, 1878)
Lycodon gibsonae G. Vogel & David, 2019
Lycodon gongshan G. Vogel & Luo, 2011
Lycodon gracilis (Günther, 1864)
Lycodon hypsirhinoides (Theobald, 1868)
Lycodon jara (Shaw, 1802)
Lycodon kundui M.A. Smith, 1943
Lycodon laoensis Günther, 1864
Lycodon liuchengchaoi Zhang et al., 2011
Lycodon meridionalis (Bourret, 1935)
Lycodon muelleri A.M.C. Duméril, Bibron & A.H.A. Duméril, 1854
Lycodon multifasciatus (Maki, 1931)
Lycodon multizonatus (Zhao & Y.-M. Jiang, 1981)
Lycodon nympha (Daudin, 1803)
Lycodon obvelatus K. Wang, Yu, G. Vogel & Che, 2020
Lycodon ophiophagus G. Vogel et al., 2009
Lycodon orientalis (Hilgendorf, 1880)
Lycodon paucifasciatus Rendahl in M.A. Smith, 1943
Lycodon philippinus Griffin, 1909
Lycodon pictus Janssen, Pham, Ngo, Le, Nguyen & Ziegler, 2019
Lycodon rosozonatus (Hu & Zhao, 1972)
Lycodon rufozonatus Cantor, 1842
Lycodon ruhstrati (Fischer, 1886)
Lycodon sealei Leviton, 1955
Lycodon semicarinatus (Cope, 1860)
Lycodon septentrionalis (Günther, 1875)
Lycodon serratus K. Wang, Yu, G. Vogel & Che, 2020
Lycodon sidiki Wostl, Hamidy, Kurniawan & E.N. Smith, 2017
Lycodon solivagus Ota & Ross, 1994
Lycodon stormi Boettger, 1892
Lycodon striatus (Shaw, 1802)
Lycodon subannulatus (A.M.C. Duméril, Bibron & A.H.A. Duméril, 1854)
Lycodon subcinctus F. Boie, 1827
Lycodon synaptor G. Vogel & David, 2010
Lycodon tessellatus Jan, 1863
Lycodon tiwarii Biswas & Sanyal, 1965
Lycodon travancoricus (Beddome, 1870)
Lycodon tristrigatus Günther, 1858
Lycodon truongi Nguyen, Duong, Wood, Grismer, 2022
Lycodon zawi Slowinski et al., 2001[6]
Lycodon zayuensis K. Jiang, Wang, Jin & Che, 2020
Lycodon zoosvictoriae Neang et al., 2014

Nota bene: A binomial authority in parentheses indicates that the species was originally described in a genus other than Lycodon.

Fitzinger LI (1826). Neue Classification der Reptilien nach ihren natürlichen Verwandtschaften. Nebst einer Verwandtschafts-tafel und einem Verzeichnisse der Reptilien-Sammlung des K.K. zoologischen Museums zu Wien. Vienna: J.G. Heubner, five unnumbered + 67 pp. + one plate. (Lycodon, new genus, p. 57). (in German and Latin).
Boulenger GA (1893). Catalogue of the Snakes in the British Museum (Natural History), Volume I., Containing the Families ... Colubridæ Aglyphæ, part.. London: Trustees of the British Museum (Natural History). (Taylor and Francis, printers). xiii + 448 pp. + Plates I-XXVIII. (Genus Lycodon, p. 348, Figure 23).
Mish, Frederick C. (Editor in Chief) (2004). Merriam-Webster's Collegiate Dictionary, Eleventh Edition. Springfield, Massachusetts: Merriam-Webster, Incorporated. 40a + 1,623 pp. ISBN 0-87779-809-5. ("lycopodium", p. 742; "odonate" p. 860).
Uetz, Peter; Hallermann, Jakob. "Lycodon ". The Reptile Database. Retrieved 11 June 2015.
"Lycodon zawi ". The Reptile Database.


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