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Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Cladus: Craniata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Classis: Reptilia
Cladus: Eureptilia
Cladus: Romeriida
Subclassis: Diapsida
Cladus: Sauria
Infraclassis: Lepidosauromorpha
Superordo: Lepidosauria
Ordo: Squamata
Subordo: Serpentes
Infraordo: Caenophidia
Superfamilia: Colubroidea

Familia: Colubridae
Subfamilia: Colubrinae
Genus: Ptyas
Species (8): P. carinata – P. dhumnades – P. dipsas – P. fusca – P. korros – P. luzonensis – P. mucosa – P. nigromarginatus

Ptyas Fitzinger, 1843: 26

Type species: Coluber blumenbachii Merrem, 1820, by original designation.

Zaocys Cope, 1861: 263 [synonymized by Taylor (1965: 738)]

Type species: Coluber dhumnades Cantor, 1842, by original designation and monotypy.
Primary references

Fitzinger, L. 1843. Systema Reptilium. Fasciculus primus. Amblyglossae. Braumüller et Seidel: Wien. vi + 106 pp. BHL Reference page.
Cope, E.D. [1860] 1861. Catalogue of the Colubridae in the Museum of the Academy of Natural Sciences of Philadelphia. Part 3. Proceedings of the Academy of Natural Sciences of Philadelphia 12: 553–566. BHL Reference page.
Meetei, A.B., Das, S., Campbell, P.D., Raha, S. & Bag, P. 2018. A study on Ptyas doriae (Boulenger, 1888) with comments on the status of Ptyas hamptoni (Boulenger, 1900) (Squamata: Colubridae: Colubrinae). Zootaxa 4457(4): 537–548. DOI: 10.11646/zootaxa.4457.4.3 Paywall Reference page.


Uetz, P. & Hallermann, J. 2021. Ptyas . The Reptile Database. Accessed on 22 May 2018.

Vernacular names
English: Asian Rat Snakes
ไทย: งูสิง

Ptyas is a genus of colubrid snakes.[1] This genus is one of several colubrid genera colloquially called "rat snakes" or "ratsnakes".

The generic name derives from Ancient Greek πτυάς, meaning "spitter", which referred to a kind of snake believed to spit venom in the eyes of humans, although in reality none of the Ptyas are known to spit venom.[2][3]

13 species are recognized:[1]

Ptyas carinata (Günther, 1858) – king korros
Ptyas dhumnades (Cantor, 1842) – Cantor's rat snake[4][5]
Ptyas dipsas (Schlegel, 1837)
Ptyas doriae Boulenger, 1888 – Doria's green snake
Ptyas fusca (Günther, 1858)
Ptyas herminae (Boettger, 1895) – Sakishima green snake
Ptyas korros (Schlegel, 1837) – Chinese ratsnake, Indo-Chinese ratsnake
Ptyas luzonensis (Günther, 1873)
Ptyas major (Günther, 1858) – Greater green snake
Ptyas mucosa (Linnaeus, 1758) – Oriental ratsnake, dhaman
Ptyas multicincta (Roux, 1907) – Many-banded green snake, North China green snake
Ptyas nigromarginata (Blyth, 1854) – green ratsnake
Ptyas semicarinata (Hallowell, 1861) – Ryukyu green snake


Ptyas at the Reptile Database. Accessed 9 March 2021.
"Ptyas dhumnades - Big-eyed Ratsnake".
"πτυάς". Academic Dictionaries and Encyclopedias.
David, P.; Vogel, G. (1996). The Snakes of Sumatra: An Annotated Checklist and Key with Natural History Notes. Edition Chimaira. Ed. Chimaira. p. 104. ISBN 978-3-930612-08-6. Retrieved 10 June 2019. "Coluber dhumnades Cantor, 1842 by original designation. Was synonymised with Ptyas by Wall (1923b) and Taylor (1965)."

Beolens, B.; Watkins, M.; Grayson, M. (2011). The Eponym Dictionary of Reptiles. The Eponym Dictionary of Reptiles. Johns Hopkins University Press. p. 162. ISBN 978-1-4214-0227-7. Retrieved 10 June 2019.

Further reading

Fitzinger, L. 1843. Systema Reptilium, Fasciculus Primus, Amblyglossae. Braumüller & Seidel. Vienna. 106 pp. + indices. (Ptyas, p. 26.)


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