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Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Cladus: Craniata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Classis: Reptilia
Cladus: Eureptilia
Cladus: Romeriida
Subclassis: Diapsida
Cladus: Sauria
Infraclassis: Lepidosauromorpha
Superordo: Lepidosauria
Ordo: Squamata
Subordo: Serpentes
Infraordo: Caenophidia
Superfamilia: Colubroidea

Familia: Colubridae
Subfamilia: Natricinae
Genus: Tropidonophis
Species: Tropidonophis aenigmaticus

Tropidonophis aenigmaticus Malnate & Underwood, 1988: 165.

Holotype: AMNH R-76633, ♂, collected by G.M. Tate & K.M. Wynn (4th Archbold Expedition) in 1953.

Type locality: "between Agamoia and Ailuluai, Fergusson Island, d’Entrecasteaux Islands, Milne Bay District, Papua New Guinea".


Amphiesma montana - Malnate, E.V., 1960: 50 (part).
Styporhynchus montanus - McDowell, S.B., 1984: 18 (part).
Styporhynchus multiscutellatus - McDowell, S.B., 1984: 19 (part).

Primary references

Malnate, E.V. & Underwood, G. 1984. Australasian Natricine Snakes of the Genus Tropidonophis. Proceedings of The Academy of Natural Sciences of Philadelphia 140(1): 59–201. JSTOR Hybrid open access journal Reference page.

Additional references

Boundy, J. 2021. Snakes of the World: A Supplement. CRC Press, Taylor and Francis Group: Boca Raton, Florida. 273 pp. ISBN 9781138618138 ebook Paywall Reference page.
Malnate, E.V. 1960. Systematic Division and Evolution of the Colubrid Snake Genus Natrix, with Comments on the Subfamily Natricinae. Proceedings of the Academy of Natural Sciences of Philadelphia 112: 41–71. JSTOR Hybrid open access journal Reference page.
McDowell, S.B. 1984. Results of the Archbold Expeditions. No. 1 12. The Snakes of the Huon Peninsula, Papua New Guinea. American Museum Novitates 2775: 1–28. (PDF) Open access Reference page.
O'Shea, M. 1996. A Guide to the Snakes of Papua New Guinea. Independent Publishing, Port Moresby, Papua New Guinea. 239 pp. Full book (PDF). Open access Reference page.
Wallach, V., Williams, K.L. & Boundy, J. 2014. Snakes of the World: A Catalogue of Living and Extinct Species. CRC Press, Taylor and Francis Group: Boca Raton, Florida. 1237 pp. ISBN 9781138034006 ebook Paywall Reference page.

Vernacular names
English: East Papuan keelback


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