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Superregnum : Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Cladus: Craniata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Superclassis/Classis: Actinopterygii
Classis/Subclassis: Actinopteri
Subclassis/Infraclassis: Neopterygii
Infraclassis: Teleostei
Megacohors: Osteoglossocephalai
Supercohors: Clupeocephala
Cohors: Euteleosteomorpha
Subcohors: Neoteleostei
Infracohors: Eurypterygia
Sectio: Ctenosquamata
Subsectio: Acanthomorphata
Divisio/Superordo: Acanthopterygii
Subdivisio: Percomorphaceae
Series: Anabantaria
Ordo: Anabantiformes
Subordo: Anabantoidei

Familia: Anabantidae
Genus: Anabas
Species: A. cobojius – A. testudineus – A. tongtruongensis

Anabas Cloquet, 1816

Type species: Perca scandens Daldorff, 1797


Coius Hamilton, 1822


Cloquet, H., 1816. [Accounts: Anabas, Anabas.] In: Dictionnaire des sciences naturelles. Strasbourg (Levrault), Paris (Le Normant). v. 2, AMA-ARGE, Supplément: 35-36. BHL
Cuvier G.L. 1816–1817. Le Règne Animal distribué d'après son organisation pour servir de base à l'histoire naturelle des animaux et d'introduction à l'anatomie comparée. Avec Figures, dessinées d'après nature. Tome II. Contenant Les reptiles, les poissons, les mollusques et les annélides. Edition 1, Deterville, Paris. pp. i–xviii + 1–532. [Pls. 9–10, in v. 4] DOI: 10.5962/bhl.title.41460 BHL Reference page.
Jayaram, K.C., 1981. The freshwater fishes of India, Pakistan, Bangladesh, Burma and Sri Lanka--a handbook. Zoological Survey of India. i-xxii + 1-475, Pls. 1-13. Online:
Kottelat, M., 2000. Notes on taxonomy, nomenclature and distribution of some fishes of Laos. Journal of South Asian Natural History v. 5 (no. 1): 83-90. Online:
Kottelat, M. 2013. The fishes of the inland waters of southeast Asia: a catalogue and core bibiography of the fishes known to occur in freshwaters, mangroves and estuaries. Raffles Bulletin of Zoology Supplement No. 27: 1–663. PDF. Reference page.
Rainboth, W.J. 1996. FAO species identification field guide for fishery purposes. Fishes of the Cambodian Mekong. Rome, FAO. 1–265, Pls. I–XXVII. Reference page.
Rao, B.V.S., 1968. Systematic studies on Anabas testudineus (Bl., 1792) and A. oligolepis Blkr., 1855. Proceedings of the Indian Academy of Sciences - Section B v. 67 (no. 5): 207-214. Online:
Roberts, T.R., 1989. The freshwater fishes of western Borneo (Kalimantan Barat, Indonesia). Memoirs of the California Academy of Sciences No. 14: i-xii + 1-210. BHL
Sen, N., 1995. Pisces. In: Fauna of Meghalaya, Part 1. Vertebrates. State Fauna Series 4. Zoological Survey of India. 483-606, Pls. 1-39, maps 1-12. Online:
Weber, M. and L. F. de Beaufort, 1922. The fishes of the Indo-Australian Archipelago. IV. Heteromi, Solenichthyes, Synentognathi, Percesoces, Labyrinthici, Microcyprini. E. J. Brill, Leiden. v. 4: i-xiii + 1-410. BHL


Anabas species list in FishBase,
Froese, R. & Pauly, D. (eds.) 2022. FishBase. World Wide Web electronic publication,, version 08/2021.
Anabas and its species (including synonyms) in Catalog of Fishes, Eschmeyer, W.N., Fricke, R. & van der Laan, R. (eds.) 2022. Catalog of Fishes electronic version.
Anabas – Taxon details on Integrated Taxonomic Information System (ITIS).
Anabas – Taxon details on National Center for Biotechnology Information (NCBI).
Anabas in the World Register of Marine Species

Vernacular names
čeština: lezoun
lietuvių: Šliaužikai
ไทย: ปลาหมอ, ปลาเข็ง, ปลาสะเด็ด
українська: Анабас

Anabas is a genus of climbing gouramies native to southern and eastern Asia. In the wild, Anabas species grow up to 30 cm (1 ft) long. They inhabit both brackish and fresh water.[2] Anabas species possess a labyrinth organ, a structure in the fish's head which allows it to breathe atmospheric oxygen, so it can be out of water for an extended period of time (6–8 hr), hence its name from the Greek anabainein ‘walk up’, from ana- ‘up’ + bainein ‘go’. They are carnivorous, living on a diet of water invertebrates and their larvae, and - in contrast to most of their relatives - are scatter spawners with no parental care. Species are found in South Asia, including India, Sri Lanka, Bangladesh, Burma, Indonesia, Malaysia, Thailand, Cambodia, and the Philippines.

There are two recognized species in the genus Anabas:[3]

Anabas cobojius (F. Hamilton, 1822) (Gangetic koi)
Anabas testudineus (Bloch, 1792) (climbing perch)


Eschmeyer, William N.; Fricke, Ron & van der Laan, Richard (eds.). "Anabas". Catalog of Fishes. California Academy of Sciences. Retrieved 11 December 2019.
Froese, Rainer and Pauly, Daniel, eds. (2012). Species of Anabas in FishBase. December 2012 version.

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