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Superregnum : Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Cladus: Craniata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Superclassis/Classis: Actinopterygii
Classis/Subclassis: Actinopteri
Subclassis/Infraclassis: Neopterygii
Infraclassis: Teleostei
Megacohors: Osteoglossocephalai
Supercohors: Clupeocephala
Cohors: Otomorpha
Subcohors: Ostariophysi
Sectio: Otophysa
Ordo: Cypriniformes
Superfamilia: Cobitoidea

Familia: Nemacheilidae
Subfamilia: Nemacheilinae
Genus: Yunnanilus
Species: Y. altus – Y. analis – Y. bailianensis – Y. bajiangensis – Y. beipanjiangensis – Y. brevis – Y. caohaiensis – Y. chui – Y. cruciatus – Y. discoloris – Y. elakatis – Y. forkicaudalis – Y. ganheensis – Y. jinxiensis – Y. longibarbatus – Y. longibulla – Y. longidorsalis – Y. macrogaster – Y. macroistainus – Y. macrolepis – Y. nanpanjiangensis – Y. niger – Y. nigromaculatus – Y. niulanensis – Y. obtusirostris – Y. pachycephalus – Y. paludosus – Y. parvus – Y. pleurotaenia – Y. pulcherrimus – Y. qujinensis – Y. sichuanensis – Y. spanisbripes – Y. tigerivinus – Y. yangzonghaiensis

Yunnanilus Nichols, 1925: 1

Type species: Nemacheilus pleurotaenia ♂ Regan, 1904. Technically type is by subsequent designation.

Eonemachilus Berg, 1938: 314, 316
Micronemacheilus Rendahl, 1944
Micronoemacheilus err. spell. -- Prokofiev, 2010
Petruichthys (subgenus of Noemacheilus) Menon, 1987: 36, 181


Nichols, J.T. 1925: Nemacheilus and related loaches in China. American Museum novitates, 171: 1–7.
Prokofiev, A. M.; 2004: A new species of the genus Micronoemacheilus Rendahl, 1944 (Balitoridae: Nemacheilinae) from Hainan (China) with notes on its status. Voprosy Ikhtiologii, 44 (3): 154–161.
Prokofiev, A. M.; 2010: Morphological classification of loaches (Nemacheilinae). Journal of Ichthyology, 50 (10): 827–913.
Zhao, Y.-H. , R. E. Gozlan & C.-G. Zhang 2011: Out of sight out of mind: current knowledge of Chinese cave fishes. Journal of fish biology, 79 (6, special issue): 1545–1562.

Yunnanilus is a genus of small stone loaches that are endemic to southeastern China, especially Guangxi and Yunnan. They are found in rivers, streams and lakes; some species are restricted to caves.[1]

There are currently 35 recognized species in this genus,[2] although other authorities cite fewer species and place a few other species formerly placed here into other genera such as Petruichthys and Micronemacheilus.[3]

Yunnanilus altus Kottelat & X. L. Chu, 1988
Yunnanilus analis J. X. Yang, 1990
Yunnanilus bailianensis J. Yang, 2013[4]
Yunnanilus bajiangensis W. X. Li, 2004
Yunnanilus beipanjiangensis W. X. Li, W. N. Mao & R. F. Sun, 1994
Yunnanilus brevis (Boulenger, 1893)
Yunnanilus caohaiensis R. H. Ding, 1992
Yunnanilus chuanheensis Jiang, Zhao, Du & Wang, 2021[5]
Yunnanilus chui J. X. Yang, 1991
Yunnanilus cruciatus (Rendahl (de), 1944)
Yunnanilus discoloris W. Zhou & J. C. He, 1989
Yunnanilus elakatis W. X. Cao & S. Q. Zhu, 1989
Yunnanilus forkicaudalis Li, 1999
Yunnanilus ganheensis L. An, B. S. Liu & W. X. Li, 2009
Yunnanilus jinxiensis Y. Zhu, L. N. Du & X. Y. Chen, 2009
Yunnanilus longibarbatus X. Gan, X. Y. Chen & J. X. Yang, 2007
Yunnanilus longibulla J. X. Yang, 1990
Yunnanilus macrogaster Kottelat & X. L. Chu, 1988
Yunnanilus macrolepis Li, Tao & Mao, 2000
Yunnanilus macrositanus W. X. Li, 1999
Yunnanilus nanpanjiangensis W. X. Li, W. N. Mao & Z. M. Lu, 1994
Yunnanilus niger Kottelat & X. L. Chu, 1988
Yunnanilus nigromaculatus (Regan, 1904) [6]
Yunnanilus niulanensis Z. M. Chen, J. Yang & J. X. Yang, 2012[7]
Yunnanilus obtusirostris J. X. Yang, 1995
Yunnanilus pachycephalus Kottelat & X. L. Chu, 1988
Yunnanilus paludosus Kottelat & X. L. Chu, 1988
Yunnanilus parvus Kottelat & X. L. Chu, 1988
Yunnanilus pleurotaenia (Regan, 1904)
Yunnanilus pulcherrimus J. X. Yang, X. Y. Chen & J. H. Lan, 2004
Yunnanilus qujinensis L. N. Du, Y. F. Lu & X. Y. Chen, 2015[6]
Yunnanilus sichuanensis R. H. Ding, 1995
Yunnanilus spanisbripes L. An, B. S. Liu & W. X. Li, 2009
Yunnanilus tigerivinus Li & Duan, 1999
Yunnanilus yangzonghaiensis Cao & Zhu, 1989


Ma, L.; and Y.-H. Zhao (2012). Cavefish of China. Pp. 107—125 in: White, W.B.; and D.C. Cuvier, editors. Encyclopedia of Caves. Elsevier. ISBN 9780123838322
Froese, Rainer and Pauly, Daniel, eds. (2017). Species of Yunnanilus in FishBase. October 2017 version.
Kottelat, M. (2012). "Conspectus_cobitidum.pdf Conspectus cobitidum: an inventory of the loaches of the world (Teleostei: Cypriniformes: Cobitoidei)" (PDF). The Raffles Bulletin of Zoology. Supplement No. 26: 1–199.
Lan, J., Gan, X., Wu, T. & Yang, J. (2013): Cave Fishes of Guangxi, China. Science Press, Beijing. 1-266.
Jiang, Wan-Sheng; Zhao, Ya-Peng; Du, Li-Na; Wang, Mo (2021-03-18). "Yunnanilus chuanheensis, a new loach species (Cypriniformes: Nemacheilidae) from the upper Lixianjiang River in Yunnan, China". Zoological Research. 42 (2): 241–245. doi:10.24272/j.issn.2095-8137.2020.287. ISSN 2095-8137. PMC 7995267. PMID 33527801.
Du, L.-N., Lu, Y.-F. & Chen, X.-Y. (2015): Yunnanilus qujinensis, a new species of loach from Yunnan, southwest China (Teleostei: Nemacheilidae). Ichthyological Exploration of Freshwaters, 26 (3): 249-254.
Chen, Z., Yang, J. & Yang, J. (2012): Description of a new species of the genus Yunnanilus Nichols, 1925 (Teleostei: Nemacheilidae) from Yunnan, China. Zootaxa, 3269: 57–64.

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