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Classification System: APG IV

Superregnum: Eukaryota
Regnum: Plantae
Cladus: Angiosperms
Cladus: Eudicots
Cladus: Core eudicots
Cladus: Rosids
Cladus: Eurosids I
Ordo: Cucurbitales

Familia: Anisophylleaceae
Genera: AnisophylleaCombretocarpusPogaPolygonanthus

Anisophylleaceae Ridl., Fl. Malay. Penin. 1: 700. (1922) as Anisophylleae

Type genus: Anisophyllea R.Br. ex Sabine, Trans. Hort. Soc. London 5: 446. (1824)


Ridley, H.N. 1922. The Flora of the Malay Peninsula 1: 700.
Chen, X., He, H. & Zhang, L.B. 2015. A monograph of the Anisophylleaceae (Cucurbitales) with description of 18 new species of Anisophyllea. Phytotaxa 229(1): 1-189. DOI: 10.11646/phytotaxa.229.1.1 Paywall Reference page.
Govaerts, R. et al. 2020. Anisophylleaceae in Kew Science Plants of the World online. The Board of Trustees of the Royal Botanic Gardens, Kew. Published online. Accessed: 2020 Apr. 18. Reference page.
International Plant Names Index. 2021. Anisophylleaceae. Published online. Accessed: Oct. 29 2021.
Schwarzbach, A.E. & Ricklefs, R.E. 2000. Systematic affinities of Rhizophoraceae and Anisophylleaceae, and intergeneric relationships within Rhizophoraceae, based on chloroplast DNA, nuclear Ribosomal DNA, and morphology. American Journal of Botany 87(4): 547–564. DOI: 10.2307/2656599 Open access Reference page.
Stevens, P.F. 2001 onwards. Angiosperm Phylogeny Website. Version 14, July 2017 [and more or less continuously updated since]. Online. Reference page.

Vernacular names
français: Anisophylléacées
中文: 异叶木科

The Anisophylleaceae are a small family with four genera and about 70 species,[1] in the order Cucurbitales, according to the APG II. However, it is more isolated from the other suprafamilal clades in this order, while it shows some similarities in flower morphology with the genus Ceratopetalum (family Cunoniaceae, order Oxalidales). Several wood features of this family are more primitive than those of the other families in the order Cucurbitales.

Previously, this family was categorized under its own order, Anisophylleales, by Takhtajan in 1997.

It is a pantropical family of shrubs and medium-sized to fairly large trees, occurring in wet, tropical forests and swamps of America, Africa and Asia.

The palmately veined leaves have a rather leathery texture, entire margins, and are often asymmetrical at the base. They have minute stipules or simply lack them. They are alternate; spiral, or distichous, or four-ranked (such as in Anisophyllea). The paired leaves may be different in size or shape.

The small flowers are regular and trimerous to pentamerous. They are usually aggregated in axillary racemes or panicles. The flower type varies considerably, most are monoecious, except Combretocarpus; which is hermaphrodite, having perfect flowers.

The inferior, tri- or quadrilocular ovary develops into a drupe or a samara (as in Combretocarpus) with usually one seed, but with three or four seeds in Poga.

Modern molecular phylogenetics suggest the following relationships:[2][3]

Cucurbitaceae (outgroup)







Christenhusz, M. J. M. & Byng, J. W. (2016). "The number of known plants species in the world and its annual increase". Phytotaxa. Magnolia Press. 261 (3): 201–217. doi:10.11646/phytotaxa.261.3.1.
Schwarzbach AE, Ricklefs RE (2000). "Systematic affinities of Rhizophoraceae and Anisophylleaceae, and intergeneric relationships within Rhizophoraceae, based on chloroplast DNA, nuclear ribosomal DNA, and morphology". Am J Bot. 87 (4): 547–564. doi:10.2307/2656599. JSTOR 2656599.
Zhang L-B, Simmons MP, Renner SS (2007). "A phylogeny of Anisophylleaceae based on six nuclear and plastid loci: Ancient disjunctions and recent dispersal between South America, Africa, and Asia" (PDF). Molecular Phylogenetics and Evolution. 44 (3): 1057–1067. doi:10.1016/j.ympev.2007.03.002. PMID 17433719.

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