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Classification System: APG IV

Superregnum: Eukaryota
Regnum: Plantae
Cladus: Angiosperms
Cladus: Eudicots
Cladus: Core eudicots
Ordo: Caryophyllales

Familia: Cactaceae
Subfamilia: Cactoideae
Tribus: Echinocereeae
Genera: ArmatocereusAustrocactusBergerocactusCarnegieaCastellanosiaCephalocereusCorryocactusDeamiaEchinocereusEscontriaEulychniaIsolatocereusJasminocereusLemaireocereusLeptocereusLophocereusLymanbensoniaMarshallocereusMitrocereusMorangayaMyrtillocactusNeoraimondiaNyctocereusPachycereusPeniocereusPfeifferaPolaskiaStenocereusStrophocactus
Accepted Natural Nothogenera: × Myrtgerocactus – × Pachebergia – × Pacherocactus – × Polascontria

Echinocereeae Buxb., Madrono 14: 179. (1958)

Type genus: Echinocereus Engelm.


Pachycereeae Buxb., Madroño 14: 186. (1958)
Type genus: Pachycereus (A.Berger) Britton & Rose
Peniocereeae Doweld Sukkulenty 4(1-2): 28. (2002)
Type genus: Peniocereus Britton & Rose
Phyllocacteae Rchb. (1841) pro parte see Hylocereeae


Buxbaum, F. 1958. Madrono 14(6): 179.
Arias Montes, S. & al. 2005. Phylogenetic relationships in Peniocereus (Cactaceae) inferred from plastid DNA sequence data. Journal of Plant Research 118(5): 317–328 (resurrecting Nyctocereus).
Arias Montes, S., Terrazas, T. & Cameron, K.M. 2003. Phylogenetic Analysis of Pachycereus (Cactaceae, Pachycereeae) based on Chloroplast and Nuclear DNA Sequences. Systematic Botany 28(3): 547–557 [1].
Barrios, D., Gonzalez-Torres, L.R., Arias, S. & Majure, L.C. 2020. Phylogeny and taxonomy of the Antillean endemic genus Leptocereus (Cactaceae) inferred from chloroplast markers and morphological evidence. Plant Systematics and Evolution 306: 1-15. DOI: 10.1007/s00606-020-01693-5 Paywall Reference page.
Sánchez, D., Arias, S. & Terrazas, T. 2014. Phylogenetic relationships in Echinocereus (Cactaceae, Cactoideae). Systematic Botany 39(4): 1183-1196. DOI: 10.1600/036364414X683831 Paywall ResearchGate Reference page.
Sánchez, D., Arias, S., Vázquez-Sánchez, M. & Terrazas, T. 2019. Combined phylogenetic analysis in Echinocereus (Cactaceae), the use of morphology, and taxonomic implications. In: Ramawat K. (eds) Biodiversity and Chemotaxonomy. Sustainable Development and Biodiversity 24: 239-269 . Springer, Cham. DOI: 10.1007/978-3-030-30746-2_11 ResearchGate Reference page.
Tapia, H.J., Bárcenas-Argüello, M.L., Terrazas, T. & Arias, S. 2017. Phylogeny and circumscription of Cephalocereus (Cactaceae) based on molecular and morphological evidence. Systematic Botany 42(4): 709-723. DOI: 10.1600/036364417X696546 Paywall ResearchGate Reference page.

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