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Classification System: APG IV

Superregnum: Eukaryota
Regnum: Plantae
Cladus: Angiosperms
Cladus: Eudicots
Cladus: Core eudicots
Cladus: Asterids
Cladus: Lamiids
Ordo: Gentianales

Familia: Rubiaceae
Subfamilia: Rubioideae
Tribus: Coussareeae
Genus: Heterophyllaea
Species: H. lycioides – H. pustulata

Heterophyllaea Hook.f., Hooker's Icon. Pl. 12: 31. (1873)

Type species: Heterophyllaea pustulata Hook.f., Hooker's Icon. Pl. 12: 31. (1873)


Lecanosperma Rusby, Bull. Torrey Bot. Club 20: 430. (1893)


Hokker, J.D. 1873. Hooker's Icones Plantarum; or figures, with brief descriptive characters and remarks of new or rare plants 12: 31. t.1134.
Govaerts, R. et al. 2020. Heterophyllaea in World Checklist of Selected Plant Families. The Board of Trustees of the Royal Botanic Gardens, Kew. Published on the internet. Accessed: 2020 Jan. 1. Reference page.
International Plant Names Index. 2020. Heterophyllaea. Published online. Accessed: Jan. 1 2020. Rubiaceae: Heterophyllaea. Missouri Botanical Garden. Published on the internet. Accessed: 2019 Dec. 17.

Heterophyllaea is a genus of two species of shrubs in the family Rubiaceae, native to Bolivia and Argentina.

Heterophyllaea pustulata also known as cegadera contains an unknown toxin that causes photosensitization in sheep and cows.[1] It can grow to between 2 and 3 meters in height and is typically found in northwest Argentina at an altitude between 2,500 and 3,000 meters above sea level.[2]


Heterophyllaea lycioides (Rusby) Sandwith
Heterophyllaea pustulata Hook.f.


"Heterophyllaea pustulata". Retrieved 2018-07-17.
C., Núñez Montoya, Susana; Núñez, Montoya, Susana C.; R., Comini, Laura; Rumie, Vittar, Belén; M., Fernández, Ivana; A., Rivarola, Viviana; L., Cabrera, José (2008). "Phototoxic effects of Heterophyllaea pustulata (Rubiaceae)". Toxicon. 51 (8): 1409–15. doi:10.1016/j.toxicon.2008.03.011. ISSN 0041-0101. PMID 18513778.

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