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Joannesia princeps - Jardim Botânico de Brasília - DSC09714

Classification System: APG IV

Superregnum: Eukaryota
Regnum: Plantae
Cladus: Angiosperms
Cladus: Eudicots
Cladus: Core eudicots
Cladus: Rosids
Cladus: Eurosids I
Ordo: Malpighiales

Familia: Euphorbiaceae
Subfamilia: Crotonoideae
Tribus: Jatropheae
Genus: Joannesia
Species: Joannesia princeps

Joannesia princeps Vell., 1798

Anda brasiliensis Raddi, Mem. Mat. Fis. Soc. Ital. Sci. Modena, Pt. Mem. Fis. 18: 404 (1820).
Anda gomesii A.Juss., Euphorb. Gen.: 39 (1824).
Andicus pentaphyllus Vell., Fl. Flumin.: 80 (1829).
Joannesia insolita Pittier, Bol. Soc. Venez. Ci. Nat. 6: 8 (1940).

Native distribution areas:

Continental: Southern America
Regional: Brazil
E. Brazil

References: Brummitt, R.K. 2001. TDWG – World Geographical Scheme for Recording Plant Distributions, 2nd Edition
Primary references

Vellozo, J.M.C., 1798. Alographia dos alkalis fixos: 199.


Govaerts, R. et al. 2020. Joannesia princeps in World Checklist of Selected Plant Families. The Board of Trustees of the Royal Botanic Gardens, Kew. Published online. Accessed: 2020 Jun 16. Reference page.
International Plant Names Index. 2020. Joannesia princeps. Published online. Accessed: Jun 16 2020.
Govaerts, R. et al. 2020. Joannesia princeps in Kew Science Plants of the World online. The Board of Trustees of the Royal Botanic Gardens, Kew. Published online. Accessed: 2020 Jun 16. Reference page. 2020. Joannesia princeps. Missouri Botanical Garden. Published online. Accessed: 16 Jun 2020.
USDA, ARS, Germplasm Resources Information Network. Joannesia princeps in the Germplasm Resources Information Network (GRIN), U.S. Department of Agriculture Agricultural Research Service. Accessed: 07-Oct-06.

Vernacular names

Joannesia princeps, the arara nut-tree or andá-açu, is a species of moderate-sized tree in the family Euphorbiaceae, with a spreading canopy, large alternate and long petioled leaves, and coarse branches. Flowers are monoecious, and fruit is a large drupe. It is endemic to east Minas Gerais, north Espírito Santo to the south of Bahia, Brazil, and threatened by habitat loss.


World Conservation Monitoring Centre (1998). "Joannesia princeps". IUCN Red List of Threatened Species. 1998: e.T32981A9740608. doi:10.2305/IUCN.UK.1998.RLTS.T32981A9740608.en. Retrieved 14 November 2021.

"Joannesia princeps". Germplasm Resources Information Network (GRIN). Agricultural Research Service (ARS), United States Department of Agriculture (USDA).
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