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Classification System: APG IV

Superregnum: Eukaryota
Regnum: Plantae
Cladus: Angiosperms
Cladus: Eudicots
Cladus: Core eudicots
Ordo: Caryophyllales

Familia: Amaranthaceae s.l.
Cladus: Chenopodiaceae s.str.
Subfamilia: Salsoloideae
Tribus: Salsoleae
Genus: Salsola
Species: Salsola paulsenii

Salsola paulsenii Litv., Izv. Turkestansk. Otd. Imp. Russk. Geogr. Obshch. 4(5): 28. 1905

Type: Uzbekistan: Turkestania: Dominium Buchara, in arenosis pr. Farab ad fl. Amu Dar'ya, 1903-09-14, N.V. Androssov & Kelow, HFR 1431. Holotype: LE; Isotypes: B, C-19022, G-G-317043/1, H, W.

named for Ove Wilhelm Paulsen


Kali paulsenii (Litv.) Akhani & E.H.Roalson, Int. J. Pl. Sci. 168(6): 946. (2007)
Heterotypic (ref. Rilke 1997, Freitag et al. 2001, Mosyakin 2017)
Salsola pellucida Litv., Sched. Herb. Fl. Ross. 8: 16 (1922)
Type: Described from the vicinity of Farab railroad station on the Amu Darya River: Buchara, pr. Farab, 1913-09-14, N. Androsov HFR 2434. Holotype: LE, Isotypes: BM000950611, C, H, K, LE.
Kali pellucidum (Litv.) Brullo, Giusso & Hrusa, Phytotaxa 201(4): 271. (2015)
(+ ref. Rilke 1999);
Salsola elegantissima Iljin, Mat. Commis. Research Exped. Acad. Sc. 26: 282. (1930)
Type: Adaevskii u., Z Ust'-Urt, Mangyshlak, Tumgacha, 1926-10-08, Rusanov 186. 2x LE .
Salsola praecox f. elegantissima Iljin, Fl. URSS 6: 216. (1936)

Native distribution areas:

Eastern Europe
E-European Russia
Middle Asia
Kazakhstan, Kirgizstan, Tadzhikistan, Turkmenistan, Uzbekistan
Armenia, Azerbaijan (incl. Nachichevan), Northern Caucasus
Western Asia
Afghanistan , Iran
Xinjiang, Gansu, Ningxia, Qinghai, Nei Mongol

Introduced in N America.

References: Brummitt, R.K. 2001. TDWG – World Geographical Scheme for Recording Plant Distributions, 2nd Edition
Primary references

Akhani, H., Edwards, G. & Roalson, E.H. 2007. Diversification of the Old World Salsoleae s.l. (Chenopodiaceae): Molecular Phylogenetic Analysis of Nuclear and Chloroplast Data Sets and a Revised Classification. International Journal of Plant Sciences 168(6): 931–956. DOI: 10.1086/518263 ResearchGate Reference page. : 946.
Brullo, C., Brullo, S., Gaskin, J.F., Giusso, G., Hrusa, G.F. & Salmeri, C. 2015. A new species of Kali (Salsoloideae, Chenopodiaceae) from Sicily, supported by molecular analysis. Phytotaxa 201(4): 256–277. (2015-03-05, epublished). DOI: 10.11646/phytotaxa.201.4.2 Reference page. : 271
Iljin, M.M. 1930. Nekotorye novye vidy flory Kazakstana. [New Species of Flora of Kazakhstan]. Mater. Kommiss. Eksped. Issl. [Mat. Commis. Research Exped. Acad. Sc.] 26: 281–285. Reference page. : 282
Iljin, M.M. 1936. Chenopodiaceae. In: Komarov, V.L. & Shishkin, B.K. (eds.): Flora URSS 6: 2–354. (russian) BHL (english translation). Reference page. : 216
Litvinov, D.I. 1905. Rasteniya beregov Aral’skago morya i Syr-Dar’inskoi oblasti, sobrannyya L.S.Bergom. Izvestiya Turkestanskago Otděla Imperatorskago Geograficheskago Obshchestva 4(5): 1–40. Reference page. : 28
Litvinov, D.I. 1922. Schedae ad Herbarium Florae Rossicae [Список растений Гербария русской флоры] 8. (Fasc. 49, 2013; bound issue with all fascicles: 1922) Reference page. : 16, no. 2434

Additional references

Mosyakin, S.L. 2017. The first record of Salsola paulsenii (Chenopodiaceae) in Ukraine, with taxonomic and nomenclatural comments on related taxa. Ukrayins'kyi Botanicnyi Zhurnal 74(5): 409–420. DOI: 10.15407/ukrbotj74.05.409 Reference page.
Mosyakin, S.L., Freitag, H. & Rilke, S. 2017. Kali versus Salsola: the instructive story of a questionable nomenclatural resurrection. Israel Journal of Plant Sciences 64: 18–30. DOI: 10.1080/07929978.2016.1256135 Reference page.
Rilke, S. 1997. Salsola paulsenii. In:
Hedge, I.C., Akhani, H., Freitag, H., Kothe-Heinrich, G., Podlech, D., Rilke, S. & Uotila, P. 1997. Chenopodiaceae. In: Rechinger, K.H. (ed.): Flora Iranica Vol. 172. 371 pp., 212 tab., Akademische Druck- u. Verlagsanstalt Graz. Reference page. : 185–187.
Rilke, S. 1999. Revision der Sektion Salsola s.l. der Gattung Salsola (Chenopodiaceae). Bibliotheca Botanica 149. Schweizerbart, Stuttgart, 189 pp. ISBN 978-3510480203. Reference page. : 155–162
Freitag, H., Hedge, I.C, Jafri, S.M.H, Kothe-Heinrich, G., Omer, S. & Uotila, P. 2001. Chenopodiaceae. In: Ali, S.I. (ed.): Flora of Pakistan 204. 217 pp., Department of Botany/Missouri Botanical Press, Karachi/St. Louis, ISBN 1-930723-10-5 eFloras. Reference page. : Salsola paulsenii.
Zhu, G., Mosyakin, S.L. & Clemants, S.E. 2004. Salsola paulsenii. In: Flora of China 5: 410.
Zhu, G., Mosyakin, S.L. & Clemants, S.E. 2004. Salsola pellucida. In: Flora of China 5: 410.


Hassler, M. 2019. Kali paulsenii. World Plants: Synonymic Checklists of the Vascular Plants of the World In: Roskovh, Y., Abucay, L., Orrell, T., Nicolson, D., Bailly, N., Kirk, P., Bourgoin, T., DeWalt, R.E., Decock, W., De Wever, A., Nieukerken, E. van, Zarucchi, J. & Penev, L., eds. 2019. Species 2000 & ITIS Catalogue of Life. Published online. Accessed: 2019 Apr. 11. Reference page.
International Plant Names Index. 2019. Salsola paulsenii. Published online. Accessed: Apr. 11 2019.
Govaerts, R. et al. 2019. Salsola paulsenii in Kew Science Plants of the World online. The Board of Trustees of the Royal Botanic Gardens, Kew. Published online. Accessed: 2019 Apr. 11. Reference page. 2019. Salsola paulsenii. Missouri Botanical Garden. Published online. Accessed: 11 Apr. 2019.

Vernacular names
қазақша: Қаңбақ
русский: Солянка Паульсена
中文: 长刺猪毛菜, chang ci zhu mao cai

Kali paulsenii[1] (Synonym Salsola paulsenii) is a species of flowering plant in the amaranth family known by the common name barbwire Russian thistle. It is native to Eurasia and it is present in the American southwest as an introduced species and sometimes a weed in sandy, disturbed habitat types. It is an annual herb forming a brambly clump of intricately branched, prostrate to erect stems growing up to a meter long. The reddish stems are lined with yellow-green, thready, fleshy, or needlelike, spine-tipped leaves a few millimeters to three centimeters long. The inflorescence is an interrupted series of flowers, with one flower per leaf axil. The flower is surrounded by a disclike array of wide, winged sepals which are whitish at the tips and pinkish at the bases.

Kali paulsenii is named for the Danish botanist and explorer Ove Paulsen.

This plant is similar to Kali tragus and sometimes forms hybrids with it.

Hossein Akhani, Gerald Edwards & Eric H. Roalson: Diversification Of The Old World Salsoleae S.L. (Chenopodiaceae): Molecular Phylogenetic Analysis Of Nuclear And Chloroplast Data Sets And A Revised Classification, in: International Journal of Plant Sciences, 168 (6), 2007, p. 931–956.

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