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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Coleopterida
Ordo: Coleoptera
Subordo: Archostemata
Superfamilia: Cupedoidea

Familia:Cupedidae
Subfamiliae (2): Cupedinae – †Mesocupedinae
Overview of Recent genera (7)

AdinolepisAscioplagaCupesParacupesPriacmaProlixocupesRhipsideigma
Name

Cupedidae Laporte, 1838

Type genus: Cupes Fabricius, 1801.

Synonymy

Cupesidae Laporte, 1838: 56

References
Primary references

Laporte [de Castelnau], F.L. 1838 ("1836"). Études entomologiques, ou descriptions d'insectes nouveaux et observations sur la synonymie [continued]. Revue Entomologique 4(1): 5–60. BHL Reference page.

Additional references

Atkins, M.D. 1963: The Cupedidae of the World. Canadian entomologist, 95(2): 140–162. [Publication date: 'February 1963', sourced from first page of article] DOI: 10.4039/Ent95140-2
Bouchard, P. & Bousquet, Y. 2020. Additions and corrections to “Family-group names in Coleoptera (Insecta)”. ZooKeys 922: 65–139. DOI: 10.3897/zookeys.922.46367 Open access. Reference page.
Bouchard, P., Bousquet, Y., Davies, A.E., Alonso-Zarazaga, M.A., Lawrence, J.F., Lyal, C.H.C., Newton, A.F., Reid, C.A.M., Schmitt, M., Ślipiński, S.A. & Smith, A.B.T. 2011. Family-group names in Coleoptera (Insecta). ZooKeys 88: 1–972. DOI: 10.3897/zookeys.88.807 Open access. PMC: 3088472 Open access. Reference page.
Kirejtshuk, A.G. 2005: A revision of the genus Cupes Fabricius, 1801 from Baltic amber and some notes on taxonomy and composition of the family Cupedidae (Coleoptera, Archostemata). Mitteilungen aus dem Geologisch-Palaeontologischen Institut der Universitaet Hamburg, 89: 55–83. page scans
Hangay, G.; Zborowski, P. 2010: A guide to the beetles of Australia. CSIRO Publishing: Collingwood VIC, Australia. ISBN 9780643094871 ISBN 0643094873
Kirejtshuk, A.G., Nel, A. & Collomb, F.-M. 2010. New Archostemata (Insecta: Coleoptera) from the French Paleocene and Early Eocene, with a note on the composition of the suborder. Annales de la Société Entomologique de France (n.s.) 46: 216–227. DOI: 10.1080/00379271.2010.10697661 Open access. Reference page.
I.Löbl & A.Smetana (eds). 2003 Catalogue of Palearctic Coleoptera. Vol. 1: Archostemata-Myxophaga-Adephaga. Apollo Books, Stenstrup, Denmark ISBN 87-88757-73-0, p. 25
Neboiss, A. 1984: Reclassification of Cupes Fabricius (s.lat.), with descriptions of new genera and species (Cupedidae: Coleoptera). Systematic entomology, 9: 443–477. DOI: 10.1111/j.1365-3113.1984.tb00520.x

Links

Australian Faunal Directory
Cupedidae in SYNOPSIS OF THE DESCRIBED COLEOPTERA OF THE WORLD
Tree of Life Web Project. 1995. Cupedidae. Version 01 January 1995 (temporary) in The Tree of Life Web Project
EoL

Vernacular names
English: reticulated beetles
한국어: 곰보벌레과
русский: Лакомки
中文: 长扁甲科

The Cupedidae are a small family of beetles, notable for the square pattern of "windows" on their elytra (hard forewings), which give the family their common name of reticulated beetles.[1]

The family consists of about 30 species in 9 genera, with a worldwide distribution. Many more extinct species are known, dating as far back as the Triassic.[1] The family Ommatidae is considered a subfamily of Cupedidae by some authors, but ommatids have been found to more closely related to Micromalthidae in genomic analysis studies.[2]

These beetles tend to be elongated with a parallel-sided body, ranging in length from 10 to 20 mm (0.39 to 0.79 in), with colors brownish, blackish, or gray. The larvae are wood-borers, typically living in fungus-infested wood, and sometimes found in wood construction.[1] The larvae eat the fungus-infested dead wood or tree roots while the adults are believed to subsist on pollen and plant sap.[3]

Males of Priacma serrata (western North America) are notable for being strongly attracted to common household bleach. This suggests that compounds in bleach may resemble attractive compounds found by the beetle in nature.[1]
Contents

1 Taxonomy
1.1 Fossil genera
2 References
3 External links

Taxonomy

Adinolepis Neboiss, 1984 Australia
Ascioplaga Neboiss, 1984 New Caledonia
Cupes Fabricius, 1801 Europe, China Paleogene, North America, Recent
Distocupes Neboiss, 1984, Australia
Paracupes Kolbe, 1898, South America
Priacma LeConte, 1874 Myanmar, Cretaceous, North America, Recent
Prolixocupes Neboiss, 1960 North America, South America
Rhipsideigma Neboiss, 1984 Madagascar, East Africa
Tenomerga Neboiss, 1984 East and Southeast Asia, New Guinea, North America, South Africa

Fossil genera

After Kirejtshuk, Nel & Kirejtshuk, 2016.[4]

†Anaglyphites Ponomarenko 1964 Karabastau Formation, Kazakhstan, Oxfordian, Shar-Teg, Mongolia, Tithonian, Hengshan Formation, China, Aptian, Dzun-Bain Formation, Mongolia, Aptian, Zaza Formation, Russia, Aptian
†"Anaglyphites" pluricavus Soriano and Delclos 2006 La Pedrera de Rúbies Formation, Spain, Barremian
†Apriacma Kirejtshuk et al. 2016 Yixian Formation, China, Aptian
†Asimma Ponomarenko 1966 Madygen Formation, Kyrgyzstan, Late Triassic
†Barbaticupes Jarzembowski et al. 2017 Burmese amber, Myanmar, Cenomanian
†Cainomerga Kirejtshuk et al. 2016 Menat Formation, France, Paleocene
†Cretomerga Kirejtshuk et al. 2016 Yixian Formation, China, Aptian
†Cupopsis Kirejtshuk et al. 2016 New Jersey amber, Turonian
†Cupidium Ponomarenko 1968 Karabastau Formation, Kazakhstan, Oxfordian
†Ensicupes Hong 1976 Dalazi Formation, Guyang Formation, China, Aptian
†Furcicupes Tan and Ren 2006 Yixian Formation, China, Aptian
†Gracilicupes Tan et al. 2006 Daohugou, China, Callovian
†Kirghizocupes Ponomarenko 1966 Madygen Formation, Kyrgyzstan, Late Triassic
†Latocupes Ren and Tan 2006 Yixian Formation, China, Aptian
†Mallecupes Jarzembowski et al. 2017 Burmese amber, Myanmar, Cenomanian
†Menatops Kirejtshuk et al. 2016 Menat Formation, France, Paleocene
†Mesocupes Martynov 1926 Karabastau Formation, Kazakhstan, Oxfordian, Las Hoyas, Spain, Barremian
†Miocupes Ponomarenko 1973 Cypris Formation, Czech Republic, Miocene
†"Platycupes" sogdianus Ponomarenko 1966 Madygen Formation, Kyrgyzstan, Late Triassic
†"Priacma" sanzii Soriano and Delclos 2006 Las Hoyas, Spain, Barremian
†Priacmopsis Ponomarenko 1966 Zaza Formation, Russia, Aptian
†Pterocupes Ponomarenko 1966 Madygen Formation, Kyrgyzstan, Late Triassic
†Taxopsis Kirejtshuk et al. 2016 Baltic amber, Eocene

References

Evans, Arthur V.; Hogue, James N. (15 November 2006). "Family Groups: Reticulated Beetles Cupedidae". Field Guide to Beetles of California. University of California Press. pp. 51–. ISBN 978-0-520-24657-7.
McKenna, Duane D.; Shin, Seunggwan; Ahrens, Dirk; Balke, Michael; Beza-Beza, Cristian; Clarke, Dave J.; Donath, Alexander; Escalona, Hermes E.; Friedrich, Frank; Letsch, Harald; Liu, Shanlin (2019-12-03). "The evolution and genomic basis of beetle diversity". Proceedings of the National Academy of Sciences. 116 (49): 24729–24737. doi:10.1073/pnas.1909655116. ISSN 0027-8424. PMC 6900523. PMID 31740605.
Jarzembowski, Edmund A.; Wang, Bo; Zheng, Daran (2017-10-01). "A new spiny reticulated beetle (Coleoptera: Cupedidae) from Cretaceous Burmese amber". Proceedings of the Geologists' Association. 128 (5): 798–802. doi:10.1016/j.pgeola.2017.07.003. ISSN 0016-7878.
Kirejtshuk, A. G.; Nel, A.; Kirejtshuk, P. A. (October 2016). "Taxonomy of the reticulate beetles of the subfamily Cupedinae (Coleoptera, Archostemata), with a review of the historical development". Invertebrate Zoology (in Russian). 13 (1): 61–190. doi:10.15298/invertzool.13.2.01. ISSN 1812-9250.

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