Embolemidae

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Hymenopterida
Ordo: Hymenoptera
Subordo: Apocrita
Superfamilia: Chrysidoidea

Familia: Embolemidae
Genera (1 + 3†): Embolemus – †Baissobius – †Cretembolemus – †Embolemopsis – Trogloembolemus –
Name

Embolemidae Förster, 1856

References

Achterberg, C. van; Kats, R.J.M. van 2000: Revision of the Palaearctic Embolemidae (Hymenoptera). Zoologische Mededelingen (Leiden), 74: 251–269.
Amarante, Roberto F. Brandao & Carpenter 1999: A new species of Embolemus Westwood from the Mata Atlantica of Brazil (Hymenoptera: Embolemidae). American Museum Novitates, 3266: 1–7.
Azevedo; Amarante 2006: New species of Embolemus (Hymenoptera, Embolemidae) from eastern Brazil. Studies on Neotropical fauna and environment, 41(2): 123–129.
Mita, T. & Olmi, M. 2018. Taxonomic additions of Embolemidae and Sclerogibbidae (Hymenoptera: Chrysidoidea) from Japan, with description of a new species of Trogloembolemus. Zootaxa 4497(4): 586–592. DOI: 10.11646/zootaxa.4497.4.8 Paywall Reference page. 
Olmi, M. 1996: A revision of the world Embolemidae (Hymenoptera Chrysidoidea). Frustula Entomologica, 18(31): 85–146.
Olmi, M. 1997: A contribution to the knowledge of the Embolemidae and Dryinidae (Hymenoptera Chrysidoidea). Bollettino di Zoologia Agraria e di Bachicoltura, 29(2): 125–150.
Olmi, M. 2004: New species of Dryinidae and Embolemidae from Madagascar (Hymenoptera Chrysidoidea). Frustula Entomologica, 25: 86–109.
Olmi, M. 2004: Embolemus kheeli: a new species of Embolemidae (Hymenoptera: Chrysidoidea) from the Dominican Republic. Zootaxa, 500: 1–6.
Olmi, M. 2004: A contribution to the knowledge of the Embolemidae of Gabon, Costa Rica and Papua New Guinea (Hymenoptera Chrysidoidea). Bollettino di Zoologia Agraria e di Bachicoltura, 36(3): 335–344.
Olmi, M. 2006: A catalogue of Dryinidae and Embolemidae of South Africa, with descriptions of new species (Hymenoptera Chrysidoidea). Frustula Entomologica, 28-29: 1–57.
Olmi, M. 2007: New Zealand Dryinidae and Embolemidae (Hymenoptera: Chrysidoidea): new records and description of Bocchus thorpei new species. Records of the Auckland Museum, 44: 5–16.
Olmi, M.; Ferrer 2003: A new species of Embolemus from Cuba (Hym., Embolemidae). Entomologist's monthly magazine, 139: 135–137.
Olmi, M.; Mita, T.; Guglielmino, A. 2014: Revision of the Embolemidae of Japan (Hymenoptera: Chrysidoidea), with description of a new genus and two new species. Zootaxa 3793(4): 423–440. DOI: 10.11646/zootaxa.3793.4.2 Reference page. 
Olmi, M.; Rasnitsyn, A.P.; Guglielmino, A. 2010: Revision of rock fossils of Dryinidae and Embolemidae (Hymenoptera: Chrysidoidea). Zootaxa, 2499: 21–38. Preview
Olmi, M.; Rasnitsyn, A.P.; Guglielmino, A. 2011: [The first record of Embolemidae (Hymenoptera Chrysidoidea) in the Rovno amber (Upper Eocene) of Ukraine: a male of Ampulicomorpha succinalis Brues.] Paleontologicheskii zhurnal, 2011(1): 66–68. [not seen, in Russian, English translation in Paleontological journal, 45(1): 73–76. (2011) DOI: 10.1134/S0031030111010138]
Ortega-Blanco, J.; Delclòs, X.; Engel, M.S. 2011: The wasp family Embolemidae in Early Cretaceous amber from Spain (Hymenoptera: Chrysidoidea). Journal of the Kansas Entomological Society, 84(1): 36–42. DOI: 10.2317/JKES100628.1
Xu, Z.-f.; He, J.-h.; Olmi, M. 2001: The Embolemidae (Hymenoptera: Chrysidoidea) from China. Entomologia Sinica, 8(3): 213–217. [1]

Links

BHL bibliography
Embolemidae – Taxon details on Encyclopedia of Life (EOL).
Global Biodiversity Information Facility. 2019. GBIF Backbone Taxonomy. Checklist dataset. Taxon: Embolemidae.
ION

Embolemidae is a family of small solitary parasitoid wasps with around 70 species in 2 genera distributed around the world.[1] The few species whose biology is known are parasites on planthopper nymphs of the families Achilidae and Cixiidae.[2] There is debate regarding the status of the genus named Ampulicomorpha by Ashmead in 1893, generally considered now to be a junior synonym of Embolemus (e.g.,[1]), though some authorities dispute this (e.g.,[2])
Biology

Females are wingless while males have wings, and in temperate regions emerge later than the females, which overwinter as adults.[3] The wingless females have been recorded from the nests of ants and small mammal burrows,[4] or under stones in pastures and grasslands, and they appear to be ant mimics. A Palearctic species, Embolemus ruddii, has been found in association with the ant species Formica fusca and Lasius flavus, while in Japan, Embolemus walkeri was taken in a nest of another ant, from the genus Myrmica.[1] A Nearctic species, Embolemus confusus, has been reared from nymphs of a planthopper in the family Achilidae, where the host fed on fungi beneath the bark of rotting logs. The wasp larva lives in a bulging sac attached to the host nymph between the second and third segments.[1][5]
Fossil species

After[2][6]

†Baissobius Rasnitsyn 1975
†Baissobius carolianus Rasnitsyn 1996 Dzun-Bain Formation, Mongolia, Early Cretaceous (Aptian)
†Baissobius minimus Rasnitsyn 1996 Zaza Formation, Russia, Aptian
†Baissobius minutus Olmi et al. 2010 Zaza Formation, Russia, Aptian
†Baissobius parvus Rasnitsyn 1975 Zaza Formation, Russia, Aptian
†Cretembolemus Olmi et al. 2014
†Cretembolemus orapensis Olmi et al. 2014 Orapa, Botswana, Late Cretaceous (Turonian)
†Embolemopsis Olmi et al. 2010
†Embolemopsis baissensis Olmi et al. 2010 Zaza Formation, Russia, Aptian
†Embolemopsis maryannae Olmi, Jarzembowski, Capradossi and Perkovsky, 2020 Wessex Formation, United Kingdom, Early Cretaceous (Barremian)
Embolemus Westwood 1833 (incl. Ampulicomorpha Ashmead, 1893)
†Embolemus antiquus Perkovsky et al., 2021 Burmese amber, Myanmar, Late Cretaceous (Cenomanian)
†Embolemus brachypterus Olmi et al., 2021 Burmese amber, Myanmar, Cenomanian
†Embolemus burmensis Perkovsky et al., 2021 Burmese amber, Myanmar, Cenomanian
†Embolemus cretacicus Perkovsky et al., 2021 Burmese amber, Myanmar, Cenomanian
†Embolemus excitus Perrichot and Engel 2011 Baltic amber, Eocene
†Embolemus janzeni (Olmi et al., 2014) Burmese amber, Myanmar, Cenomanian
†Embolemus micropterus Olmi et al., 2021 Burmese amber, Myanmar, Cenomanian
†Embolemus ohmkuhnlei Perkovsky et al., 2021 Burmese amber, Myanmar, Cenomanian
†Embolemus perialla (Ortega-Blanco et al., 2011) Spanish amber, Early Cretaceous (Albian)
†Embolemus quesnoyensis (Chény, Guillam, Nel and Perrichot, 2020) Oise amber, France, Eocene (Ypresian)
†Embolemus succinalis (Brues, 1933) Baltic amber, Rovno amber, Eocene
†Embolemus zherikhini Perkovsky et al., 2021 Burmese amber, Myanmar, Cenomanian
†Ponomarenkoa Olmi 2010
†Ponomarenkoa burmensis Perkovsky et al., 2021 Burmese amber, Myanmar, Cenomanian
†Ponomarenkoa ellenbergeri Olmi et al. 2013 Burmese amber, Myanmar, Cenomanian
†Ponomarenkoa polonica Ponomarenko 1988 Baltic amber, Eocene

References

van Achterberg, Cornelis & Kats, R.. (2000). Revision of the Palaearctic Embolemidae (Hymenoptera). Zoöl. Med. 74 (2000), 17: 251-269.
Perkovsky, Evgeny E.; Olmi, Massimo; Müller, Patrick; Guglielmino, Adalgisa; Jarzembowski, Edmund A.; Capradossi, Leonardo; Rasnitsyn, Alexandr P. (November 2020). "A review of the fossil Embolemidae (Hymenoptera: Chrysidoidea), with description of seven new species and history of the family". Cretaceous Research: 104708. doi:10.1016/j.cretres.2020.104708.
J.T. Burn (1997). "Embolemus ruddii (Westwood,1833)". Bees, Wasps and Ants Recording Society. Retrieved 8 July 2017.
Simon van Noort (2017). "Embolemidae". WaspWeb: Hymenoptera of the Afrotropical region. Iziko Museums of South Africa. Retrieved 8 July 2017.
"HYMENOPTERA, Embolemidae (Chrysidoidea) (formerly Bethyloidea)". University of California, Riverside. Retrieved 8 July 2017.
Ross, Andrew (21 June 2021). "Burmese (Myanmar) amber taxa, on-line supplement v.2021.1" (PDF). National Museums Scotland. Retrieved 5 November 2021.