Hydroptilidae

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Panorpida
Cladus: Amphiesmenoptera
Ordo: Trichoptera
Subordo: Annulipalpia
Superfamilia: Rhyacophiloidea

Familia:Hydroptilidae
Subfamilia: Hydroptilinae
Tribus: Hydroptilini
Genus: Oxyethira
Subgenera (3): O. (Pacificotrichia) – O. (Trichoglene) –

Species (223):
Directory A·B·C·D·E·F·G·H·I·J·K·L·M·N·O·P·Q·R·S·T·U·V·W·X·Y·Z
A

O. abacatia – O. absona – O. acegua – O. aculea – O. acuta – O. aeola – O. ahipara – O. alaluz – O. albaeaquae – O. albiceps – O. allagashensis – O. anabola – O. andina – O. angustella – O. apinolada – O. araya – O. archaica – O. arctodactyla – O. argentinensis – O. arizona – O. arok – O. artuvillosus – O. aspera – O. assia – O. azteca –
B

O. bamaga – O. baritu – O. bettyae – O. bicornuta – O. bidentata – O. bifurcata – O. bogambara – O. boreella – O. braziliensis – O. brevis – O. burkina –
C

O. caledoniensis – O. calori – O. campanula – O. campesina – O. chrysocara – O. circaverna – O. coercens – O. colombiensis – O. columba – O. complicata – O. copina – O. cornuta – O. costaricensis – O. cotula – O. cuernuda – O. culebra –
D

O. dactylonedys – O. dalmeria – O. datra – O. delcourti – O. derek – O. desadorna – O. diplospissa – O. discaelata – O. distinctella – O. dorsennus – O. driesseni – O. dualis – O. dunbartonensis –
E

O. ecornuta – O. efatensis – O. elerobi – O. elora – O. espinada – O. espirita – O. ezoensis –
F

O. falcata – O. fijiensis – O. flagellata – O. flavicornis – O. florida – O. forcipata – O. frici –
G

O. galekoluma – O. garifosa – O. geminata – O. glasa – O. gomera – O. gracilianoi – O. grisea – O. guariba –
H

O. hainanensis – O. harpagella – O. harpeodes – O. hartigi – O. hilosa – O. hozosa – O. hyalina –
I

O. iannuzzae – O. iglesiasi – O. ikal – O. inaequispina – O. incana – O. indorsennus – O. insularis – O. itascae –
J

O. jamaicensis – O. janella – O. josifovi –
K

O. kelleyi – O. kingi – O. kirikiriroa – O. klingstedti –
L

O. lagunita – O. laodameia – O. lobophora – O. longipenis – O. longispinosa – O. longissima – O. luanae – O. lumipollex – O. lumosa –
M

O. macrosterna – O. maranhensis – O. maryae – O. matadero – O. maya – O. mcgregori – O. melasma – O. merga – O. michiganensis – O. mienica – O. minima – O. mirabilis – O. mirebalina – O. misionensis – O. mithi – O. mocoi –
N

O. novasota –
O

O. obscura – O. obtatus – O. orellanai – O. oropedion – O. ortizorum –
P

O. paieon – O. pallida – O. paludicola – O. paramartha – O. parazteca – O. parce – O. paritentacula – O. peruviana – O. pescadori – O. petei – O. picita – O. ping – O. pirisinui – O. plumosa – O. poapi – O. presilla – O. pseudofalcata – O. puertoricensis –
Q

O. quinquaginta – O. quiramae –
R

O. rachanee – O. rafaeli – O. ramosa – O. rareza – O. redunca – O. retracta – O. retrosa – O. ritae – O. rivicola – O. roberti – O. rossi –
S

O. sagittifera – O. santiagensis – O. savanniensis – O. scaeodactyla – O. scopulina – O. scutica – O. sechellensis – O. sencilla – O. septentrionalis – O. serrata – O. setosa – O. sichuanensis – O. sida – O. sierruca – O. simplex – O. simulatrix – O. singularis – O. sininsigne – O. sinistra – O. smolpela – O. spinosella – O. spirogyrae – O. spissa –
T

O. tamandua – O. tamperensis – O. tasmaniensis – O. tega – O. teixeirai – O. tenei – O. tenuella – O. tica – O. tiunovae – O. tompa – O. torquata – O. torresiana – O. touba – O. triangulata – O. tristella – O. tropis –
U

O. ulmeri – O. una – O. unidentata – O. unispina –
V

O. vaina – O. velocipes – O. verna – O. vipera – O. volsella –
W

O. waipoua – O. warramunga –
Z

O. zeronia – O. zilaba

[source: & Johanson, 2015flife.org/annual-checklist/2012/browse/tree/id/2366491 Catalogue of Life: 2012 Annual Checklist, plus (26) O. arok – O. bettyae – O. chrysocara – O. derek – O. driesseni – O. elora – O. espirita – O. geminata – O. kirikiriroa – O. laodameia – O. longipenis – O. luanae – O. mcgregori – O. paieon – O. paludicola – O. pescadori – O. ping – O. poapi – O. quiramae – O. redunca – O. scopulina – O. sinistra – O. tamandua – O. tiunovae – O. tompa – O. torquata
]

Add (16) with subgenera (in brackets): O. digitata (Pacificotrichia) – O. enigmatica (Pacificotrichia) – O. mouirange (Pacificotrichia) – O. nehoue (Pacificotrichia) – O. ouenghi (Pacificotrichia) – O. quadrata (Pacificotrichia) – O. rougensis (Pacificotrichia) – O. spicula (Pacificotrichia) – O. abbreviata (Trichoglene) – O. amieu (Trichoglene) – O. houailou (Trichoglene) – O. incurvata (Trichoglene) – O. parinsularis (Trichoglene) – O. perignonica (Trichoglene) – O. spinifera (Trichoglene) – O. tiwaka (Trichoglene) –

Not assignable to subgenus (1): O. macropennis –
Name

Oxyethira Eaton, 1873

Synonyms

Dactylotrichia
Dampfitrichia
Holarctotrichia
Kelleyella
Loxotrichia
Mesotrichia
Oxytrichia
Tanytrichia

References

Eaton 1873: Trans. ent. Soc. London, 1873, 143.
Flint, O.S. 1991: Studies of Neotropical Caddisflies, XLV: The Taxonomy, Phenology, and Faunistics of the Trichoptera of Antioquia, Colombia. Smithsonian Contributions to Zoology, 520: 1–113. full article. Reference page.
Mey, W., 1998: Contribution to the knowledge of the caddisflies of the Philippines 2. The species of the Mt. Agtuuganon Range on Mindanao (Insecta: Trichoptera). Nachrichten des Entomologischen Vereins Apollo Supplement 17: 537–576.
de Souza, W.R.M. & Santos, A.P.M. 2017. Taxonomic study of the genus Oxyethira Eaton 1873 (Trichoptera: Hydroptilidae) from Northeast Brazil: Eleven new species and distributional records. Zootaxa 4236(3): 484–506. DOI: 10.11646/zootaxa.4236.3.4. Reference page.
Wells, A. & Dostine, P. 2016. New and newly recorded micro-caddisfly species (Insecta: Trichoptera: Hydroptilidae) from Australia’s north, including islands of Torres Strait. Zootaxa 4127(3): 591–600. DOI: 10.11646/zootaxa.4127.3.11. Full article (PDF). Reference page.
Wells, A. & Johanson, K.A. 2015. Review of New Caledonian species of Oxyethira Eaton, with description of 17 new species, and new records for Hydroptila Dalman and Hellyethira Neboiss (Trichoptera, Hydroptilidae). Zookeys 530:Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Panorpida
Cladus: Amphiesmenoptera
Ordo: Trichoptera
Subordo: Annulipalpia
Superfamilia: Rhyacophiloidea

Familia:Hydroptilidae
Subfamiliae: Hydroptilinae – Ptilocolepinae

Overview of genera: Allotrichia – Pseudoxyethira
Name

Hydroptilidae Stephens, 1836
References

Stephens, J.F. 1836. Family II—Hydroptilidae mihi. Illustrations of British entomology; or, A synopsis of indigenous insects: containing their generic and specific distinctions. v. 10 (Mandibulata v. 6) 151–153. BHL Reference page.
Angrisano, E.B. & Sganga, J.V. 2009. New species of Hydroptilidae (Trichoptera) from Salto Encantado Provincial Park (Misiones province, Argentina). Zootaxa 2162: 57–68. Abstract & excerpt (PDF).
Ito, T. 2017. The genus Pseudoxyethira Schmid (Trichoptera, Hydroptilidae) in Japan. Zootaxa 4319(1): 194–200. DOI: 10.11646/zootaxa.4319.1.12. Reference page.
Parys, K.A. & Harris, S.C. 2013. Larva of Nothotrichia shasta Harris & Armitage (Trichoptera: Hydroptilidae) from California, USA, with its phylogenetic and taxonomic implications. Zootaxa 3620(4): 589–595. DOI: 10.11646/zootaxa.3620.4.8 Reference page.
De Souza, W.R.M., Santos, A.P.M., Lima, L.R.C & Pinheiro, U. 2013. A new species and new records of microcaddisflies (Trichoptera: Hydroptilidae) from northeastern Brazil. Zootaxa 3700(4): 583–587. DOI: 10.11646/zootaxa.3700.4.6 Reference page.
Souza, W.R.M. De, Santos, A.P.M. & Takiya, D.M. 2016. Three new species of Stactobiinae (Trichoptera: Hydroptilidae) with the first record of Orinocotrichia Harris, Flint & Holzenthal from Brazil; in: Pinto et al., Living among dragons: A festschrift in honor to Dr. Angelo B. M. Machado on occasion of his 80th birthday. Zootaxa 4078(1): 337–343. DOI: 10.11646/zootaxa.4078.1.28 Full article (PDF).Reference page.
Thomson, R.E., Frandsen, P.B. & Holzenthal, R.W. 2022. A preliminary molecular phylogeny of the family Hydroptilidae (Trichoptera): an exploration of combined targeted enrichment data and legacy sequence data. Pp 467–488 In Pauls, S.U., Thomson, R. & Rázuri-Gonzales, E. (eds.). Special Issue in Honor of Ralph W. Holzenthal for a Lifelong Contribution to Trichoptera Systematics Zookeys 1111ː 1–488. Reference page. . DOI: 10.3897/zookeys.1111.85361 Open access Reference page.
Wells, A. 2012. New synonyms in the Australian micro-caddisfly fauna (Trichoptera: Hydroptilidae). Zootaxa 3177: 66–68. Preview PDF

Vernacular names
中文: 小石蛾科

37–90. DOI: 10.3897/zookeys.530.6047 Preview (PDF) Reference page.

The Hydroptilidae are a large family of caddisflies (Trichoptera) with a worldwide distribution. They are commonly known as microcaddisflies or purse-case caddisflies, in reference to two characteristic traits of this family: Hydroptilidae are much smaller than other caddisflies, rarely exceeding 5 mm (0.20 in) in length. Their larvae do not build a protective case until the final instar of their growth. At that time however, they build a typically Purse-shaped case, either portable or stuck to the substrate, in which the larva finishes growth and pupates.[1]
Contents

1 Systematics and taxonomy
2 Genera
3 References
4 External links

Systematics and taxonomy

Their systematic placement among the caddisflies is still disputed. They are traditionally placed in the suborder Spicipalpia, which do not seem to be a natural monophyletic group, but rather an evolutionary grade of moderately advanced caddisflies. Some authors downrank the Spicipalpia to a superfamily of the more basal Annulipalpia and call them Rhyacophiloidea (which otherwise refers to a subfamily of suborder Spicipalpia), but recent studies generally rejected this view.[1]

More often, the Hydroptilidae are placed in a monotypic superfamily Hydroptiloidea, either in the Spicipalpia or – probably more appropriately considering the present state of caddisfly phylogeny – incertae sedis in the Trichoptera. It may be that the Glossosomatoidea are particularly closely related to the Hydroptilidae; together they might even be closer to the most advanced caddisflies (the tube case caddisflies, Integripalpia) than any other living caddisfly. But this view is almost as disputed as including the Spicipalpia in the Annulipalpia. In any case, were Glossosomatoidea and Hydroptiloidea to be merged into a single superfamily, the older name Hydroptiloidea would apply for the combined group.[1]

The peculiarly apomorphic genera Palaeagapetus and Ptilocolepus have been separated as subfamily Ptilocolepinae, while all other genera form the Hydroptilinae family with its multiple tribes. It is far from certain that the Ptilocolepinae are the living fossils such an arrangement would imply them to be.
Genera

Around 70 genera with at least 1,700 species have been described from this family:[2]

Hydroptilinae Stephens, 1836

Acanthotrichia Wells, 1982
Acritoptila Wells, 1982
Agraylea Curtis, 1834
Allotrichia McLachlan, 1880
Austratrichia Wells, 1982
Cyclopsiella Kjaerandsen, 1997
Dhatrichia Mosely, 1948
Hellyethira Neboiss, 1977
Hydroptila Dalman, 1819
Hydroptilina Martynov, 1934
Jabitrichia Wells, 1990
Microptila Ris, 1897
Missitrichia Wells, 1991
Mulgravia Wells, 1982
Oxyethira Eaton, 1873
Paroxyethira Mosely, 1924
Paucicalcaria Mathis & Bowles, 1989
Tangatrichia Wells & Andersen, 1995
Tricholeiochiton Kloet & Hincks, 1944
Ugandatrichia Mosely, 1939
Vietrichia Olah, 1989
Wlitrichia Kjaerandsen, 1997
Xuthotrichia Mosely, 1934

Leucotrichiinae Flint, 1970

Abtrichia Mosely, 1939
Acostatrichia Mosely, 1939
Alisotrichia Flint, 1964
Anchitrichia Flint, 1970
Ascotrichia Flint, 1983
Betrichia Mosely, 1939
Celaenotrichia Mosely, 1934
Cerasmatrichia Flint, Harris & Botosaneanu, 1994
Ceratotrichia Flint, 1992
Costatrichia Mosely, 1937
Eutonella Mueller, 1921
Leucotrichia Mosely, 1934
Mejicanotrichia Harris & Holzenthal, 1997
Peltopsyche Mueller, 1879
Scelobotrichia Harris & Bueno-Soria, 1993
Zumatrichia Mosely, 1937

Neotrichiinae Ross, 1956

Kumanskiella Harris & Flint, 1992
Mayatrichia Mosely, 1937
Neotrichia Morton, 1905
Taraxitrichia Flint & Harris, 1991

Ochrotrichiinae Marshall, 1979

Angrisanoia Özdikmen, 2008
Caledonotrichia Sykora, 1967
Maydenoptila Neboiss, 1977
Metrichia Ross, 1938
Ochrotrichia Mosely, 1934
Nothotrichia Flint, 1967
Ragatrichia Oláh & Johanson, 2011
Rhyacopsyche Mueller, 1879

Orthotrichiinae Nielsen, 1948

Ithytrichia Eaton, 1873
Orthotrichia Eaton, 1873

Stactobiinae Botosaneanu, 1956

Bredinia Flint, 1968
Byrsopteryx Flint, 1981
Catoxyethira Ulmer, 1912
Chrysotrichia Schmid, 1958
Flintiella Angrisano, 1995
Niuginitrichia Wells, 1990
Orinocotrichia Harris, Flint & Holzenthal, 2002
Parastactobia Schmid, 1958
Plethus Hagen, 1887
Scelotrichia Ulmer, 1951
Stactobia McLachlan, 1880
Stactobiella Martynov, 1924
Tizatetrichia Harris, Flint & Holzenthal, 2002

incertae sedis

†Burminoptila Botosaneanu, 1981
Dibusa Ross, 1939
Dicaminus Mueller, 1879
†Electrotrichia Ulmer, 1912
Macrostactobia Schmid, 1958
†Novajerseya Botosaneanu, Johnson & P. R. Dillon, 1998
Orphninotrichia Mosely, 1934

References

Ralph W. Holzenthal; Roger J. Blahnik; Aysha Prather; Karl Kjer (January 12, 2010). "Trichoptera". Tree of Life Web Project. Retrieved April 4, 2010.
"Classification Hierarchy". Trichoptera World Checklist. Clemson University. Archived from the original on June 10, 2010. Retrieved April 4, 2010.