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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Coleopterida
Ordo: Coleoptera
Subordo: Polyphaga
Infraordo: Elateriformia
Superfamilia: Scirtoidea

Familia: Scirtidae
Subfamiliae (3): Nipponocyphoninae - Scirtinae - Stenocyphoninae

[source: Lawrence & Yoshitomi (2007: 522–523)]

Check (1): †Mesernobiinae
Overview of genera (65 + 8†)

AccolabassAmplectopus – Anticyphon – AtopidaAustrocyphonBiroguineusBrachycyphonByrrhopsisCalvariumChameloscyphonChilarboreus – Contacyphon – Cygnocyphon – Cyphanodes – Cyphanus – Cyphotelus – Cyprobius – Daploeuros – Dasyscyphon – Dermestocyphon – Elodes – Eurycyphon – Exagontus – Eximiocyphon – Exneria – Exochomoscirtes – Herthania – Heterocyphon – Hydrocyphon – Leptocyphon – Macrocyphon – Macrodascillus – Macrohelodes – Memorocyphon – Mescirtes – Mesocyphon – Microcara – Nanocyphon – Nipponocyphon – Nothocyphon – Nyholmia – Odeles – Ora – Pachycyphon – Papuacyphon – Paracyphon – Parelodes – Peneveronatus – Petrocyphon – Prionocara – Prionocyphon – Prionoscirtes – Pseudomicrocara – Pseudoscirtes – Sacodes – Sarabandus – Scirtes – Spaniosdascillus – Stenactyla – Stenocyphon – Tasmanocyphon – Tectocyphon – Veronatus – Yoshitomia – Ypsiloncyphon – ?Brounicyphon – †Actinocerites – †Brachelodes – †Brachypodites – †Cyphonogenius – †Expansifemoria – †Miocyphon – †Plagiocyphon – †Sinodryopites
Name

Scirtidae Fleming, 1821

Synonyms

Cyphonidae Stephens, 1829
Elodidae Shuckard, 1840
Helodidae Agassiz, 1846
†Sinodryopitidae Hong, 2002 [syn. Kirejtshuk & Azar (2008: 29)]

References

Cooper, S.J.B., Watts, C.H.S., Saint, K.M. & Leijs, R. 2014: Phylogenetic relationships of Australian Scirtidae (Coleoptera) based on mitochondrial and nuclear sequences. Invertebrate systematics 28(6): 628–642. DOI: 10.1071/IS13046 Reference page.
Heuss, K. 2008: Zur Taxonomie und Ökologie der Schlammkäfer des Baltischen Bernsteins mit einem Gattungsschlüssel und der Beschreibung einer neuen Art (Coleoptera: Scirtidae) [The taxonomy and ecology of the marsh beetles of the Baltic amber with key to genera and description of a new species (Coleoptera: Scirtidae).] Entomologische Zeitschrift, 118(3): 131–134. PDF
Jäch, M.A.; Madl, M. 2009: Water beetles of Seychelles (Coleoptera). Pp. 11-35, plates 1-2 in: Gerlach, J. (ed.) The Coleoptera of the Seychelles Islands. Pensoft Publishers, Sofia-Moscow. ISBN 9789546424983
Kiałka, A. & Ruta, R. 2017. An illustrated catalogue of the New Zealand marsh beetles (Coleoptera: Scirtidae). Zootaxa 4366(1): 1–76. DOI: 10.11646/zootaxa.4366.1.1. Reference page.
Kirejtshuk, A.G. & Azar, D. 2008. New taxa of beetles (Insecta, Coleoptera) from Lebanese amber with evolutionary and systematic comments. Alavesia 2: 15–46. Full article (PDF) Reference page.
Klausnitzer, B. 2004: New species of Scirtidae (Coleoptera) from Baltic amber (Part 1). Entomologische Nachrichten und Berichte, 48(2): 99–103.
Klausnitzer, B. 2008: Insecta: Coleoptera: Scirtidae. Süßwasserfauna von Mitteleuropa, 20/17. Spektrum. Google books
Klausnitzer, B. 2009: Beschreibung einer neuen Gattung der Familie Scirtidae aus der Orientalischen Region (Coleoptera) (144. Beitrag zur Kenntnis der Scirtidae). Beiträge zur entomologie, 59(1): 247–261.
Klausnitzer, B. 2011: New species of Scirtidae (Coleoptera) from Baltic amber (Part 2). Entomologische Zeitschrift, 121(1): ?-?
Lawrence, J.F. 2005: 15.4. Scirtidae Fleming, 1821. Pp. 443-450 in: Beutel, R.G. & Leschen, R.A.B. (volume eds.) Coleoptera, Beetles. Volume 1: Morphology and systematics (Archostemata, Adephaga, Myxophaga, Polyphaga partim). In: Kristensen, N.P. & Beutel, R.G. (eds.) Handbook of zoology. A natural history of the phyla of the animal kingdom. Volume IV. Arthropoda: Insecta. Part 38. Berlin, New York: Walter de Gruyter.
Lawrence, J.F. & Yoshitomi, H. 2007. Nipponocyphon, a new genus of Japanese Scirtidae (Coleoptera) and its phylogenetic significance. Elytra 35(2): 507–527. Full issue. Reference page.
Ruta, R. 2009: Revision of Scirtidae (Insecta: Coleoptera) described by Victor Ivanovitsch Motschulsky. Zootaxa, 2210: 26–50. Abstract & excerpt
Ruta, R. 2016. Anticyphon gen. nov., a new genus of Scirtidae (Coleoptera: Scirtoidea) inhabiting high altitude Andean cloud forests and páramo formation. Zootaxa 4175(4): 301–318. DOI: 10.11646/zootaxa.4175.4.1. Reference page.
Ruta, R. 2013. Review of Scirtidae (Coleoptera: Scirtoidea) described by Johan Christian Fabricius (1745–1808). Zootaxa 3646(1): 51–67. DOI: 10.11646/zootaxa.3646.1.4 Reference page.
Ruta, R. & Libonatti, M.L. 2016. Redescriptions of Scirtidae (Coleoptera: Scirtoidea) described by Carl Henrik Boheman (1796–1868) with notes on Scirtes adustus diversenotatus Pic, 1930. Zootaxa 4072(2): 203–216. DOI: 10.11646/zootaxa.4072.2.4. Reference page.
Yoshitomi, H. 2005: Systematic revision of the family Scirtidae of Japan, with phylogeny, morphology and bionomics (Insecta: Coleoptera, Scirtoidea). Japanese journal of systematic entomology (monographic series), (3)
Yoshitomi, H.; Ruta, R. 2010: SCIRTIDAE: Checklist of the taxa recorded from Melanesia (Moluccas to Fiji) (Coleoptera). Pp. 401-402 in: Jäch, M.A. & Balke, M. (eds.): Water beetles of New Caledonia (part 1). Monographs on Coleoptera, 3 PDF available here
Catalogue of Palearctic Coleoptera. Vol. 3 ed. I. Lobl, & A. Smetana, Apollo Books, Stenstrup, Denmark, 2006
ISBN 87-88757-59-5, p.316
Zwick, P. 2015: Australian Marsh Beetles (Coleoptera: Scirtidae). 8. The new genera Cygnocyphon, Eximiocyphon, Paracyphon, Leptocyphon, Tectocyphon, and additions to Contacyphon de Gozis, Nanocyphon Zwick and Eurycyphon Watts. Zootaxa 3981(4): 451–490. DOI: 10.11646/zootaxa.3981.4.1. Preview (PDF) Reference page.

Links

Australian Faunal Directory
Scirtidae in SYNOPSIS OF THE DESCRIBED COLEOPTERA OF THE WORLD
Tree of Life Web Project. 2007. Scirtidae. Marsh beetles. Version 25 May 2007 (temporary) in The Tree of Life Web Project
Atlas of the family Scirtidae of Russia
EoL

Vernacular names
English: marsh beetles
latviešu: Dumbraiņu dzimta
norsk: Hårbiller
русский: Трясинники

Scirtidae is a family of beetles (Coleoptera). These beetles are commonly referred to as marsh beetles, as the larvae are typically associated with stagnant water, but can be found in flowing water. Adults prefer decomposing plant material near the water's edge.[2] More than an estimated 600 species are known worldwide, distributed among at least 60 genera.[3]

Genera

These 81 genera belong to the family Scirtidae:[1][4][5][6]

Accolabass Watts, 2009
Alpestriscyphon Watts, Cooper & Libonatti, 2020
Amplectopus Sharp, 1886
Anocyphon Watts, Cooper & Libonatti, 2020
Anthocara Watts, Cooper & Libonatti, 2020
Anticyphon Ruta, 2016
Atopida White, 1846
Austrocyphon Zwick, 2013
Brachelodes Yablokov-Khnzorian, 1961
Brachycyphon Fairmaire, 1896
Byrrhopsis Champion, 1913
Calvariopsis Ruta, 2019
Calvarium Pic, 1918
Chameloscyphon Watts, 2011
Chilarboreus Ruta, 2011
Contacyphon Des Gozis, 1886
Copiacyphon Watts, Cooper & Libonatti, 2020
Cygnocyphon Zwick, 2015
Cyphanodes Broun, 1893
Cyphanus Sharp, 1878
Cyphon Paykull, 1799
Cyphonogenius Yablokov-Khnzorian, 1961
Cyphotelus Sharp, 1878
Cyprobius Sharp, 1878
Daploeuros Watts, 2011
Dasyscyphon Watts, 2011
Elodes Latreille, 1796
Eurycyphon Watts, 2011
Exneria Klausnitzer, 2013
Exochomoscirtes Pic, 1916
Furcacyphon Watts, Cooper & Libonatti, 2020
Herthania Klausnitzer, 2006
Heterocyphon Armstrong, 1953
Hydrocyphon Redtenbacher, 1858
Indiocyphon Pic, 1918
Latuscara Watts, Cooper & Libonatti, 2020
Leptocyphon Zwick, 2015
Macrocyphon Pic, 1918
Macrodascillus Carter, 1935
Meatopida Kialka & Ruta, 2018
Memorocyphon Pic, 1918
Mescirtes Motschulsky, 1863
Mesocyphon Sharp, 1878
Microcara Thomson, 1859
Miocyphon Wickham, 1914
Nanocyphon Zwick, 2013
Nasutuscyphon Watts, Cooper & Libonatti, 2020
Nektriscyphon Watts, Cooper & Libonatti, 2020
Nipponocyphon Lawrence & Yoshitomi, 2007
Nothocyphon Zwick, 2015
Nyholmia Klausnitzer, 2013
Odeles Klausnitzer, 2004
Ora Clark, 1865 (flea marsh beetles)
Papuacyphon Zwick, 2014
Paracyphon Zwick, 2015
Peneveronatus Armstrong, 1953
Perplexacara Watts, Bradford & Cooper, 2021
Petrocyphon Watts, 2011
Pictacara Watts, Cooper & Libonatti, 2020
Plagiocyphon Yablokov-Khnzorian, 1961
Prionocyphon Redtenbacher, 1858
Prionoscirtes Champion, 1897
Pseudomicrocara Armstrong, 1953
Pumiliocara Watts, Cooper & Libonatti, 2020
Ruborcara Watts, Cooper & Libonatti, 2020
Sacodes LeConte, 1853
Saltuscyphon Watts, Cooper & Libonatti, 2020
Saprocyphon Watts, Cooper & Libonatti, 2020
Sarabandus Leech, 1955
Scirtes Illiger, 1807
Sisyracyphon Watts, Cooper & Libonatti, 2020
Spaniosdascillus Watts, 2011
Spilotocyphon Watts, Cooper & Libonatti, 2020
Stenocyphon Lawrence, 2001
Tasmanocyphon Zwick, 2013
Tectocyphon Zwick, 2015
Tenebriocyphon Watts, Cooper & Libonatti, 2020
Vadumcyphon Watts, Cooper & Libonatti, 2020
Veronatus Sharp, 1878
Yoshitomia Klausnitzer, 2013
Ypsiloncyphon Klausnitzer, 2009

Fossil genera

†Mesernobius Engel 2010 Burmese amber, Myanmar, Cenomanian
†Miocyphon Wickham 1914 Florissant Formation, Colorado, Eocene

References

"Scirtidae Report". Integrated Taxonomic Information System. Retrieved 2018-04-28.
Epler, J.H. (2010). The Water Beetles of Florida – an identification manual for the families Chrysomelidae, Curculionidae, Dryopidae, Dytiscidae, Elmidae, Gyrinidae, Haliplidae, Helophoridae, Hydraenidae, Hydrochidae, Hydrophilidae, Noteridae, Psephenidae, Ptilodactylidae and Scirtidae. Tallahassee: Florida Department of Environmental Protection. p. 414.
Thomas, Michael C. (2002). Arnett, Ross H.; Thomas, M. C.; Skelley, P. E.; et al. (eds.). American Beetles, Volume II: Polyphaga: Scarabaeoidea through Curculionoidea. Boca Raton: CRC Press. ISBN 0-8493-0954-9.
"Browse Scirtidae". Catalogue of Life. Retrieved 2018-04-28.
"Scirtidae". GBIF. Retrieved 2018-04-28.
"Scirtidae Family Information". BugGuide.net. Retrieved 2018-04-28.

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