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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Cladus: Pancrustacea
Cladus: Allotriocarida
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Holometabola
Superordo: Hymenopterida
Ordo: Hymenoptera
Subordo: Apocrita
Superfamilia: Proctotrupoidea

Familia: Heloridae
Subfamiliae (1 + 1†): Helorinae - †Protohelorinae
Overview of genera
†Archaeohelorus – ...

Heloridae is a family of wasps in the order Hymenoptera known primarily from fossils, and only one extant genus, Helorus, with 12 species found worldwide.[1][2][3][4][5] Members of Helorus are parasitic on green lacewings.[1]
Extant genera

Helorus (12 spp. worldwide)

Fossil genera

†Archaeohelorus Daohugou, China, Middle Jurassic (Callovian)
†Bellohelorus Li et al. 2017 Yixian Formation, China, Early Cretaceous (Aptian)
†Conohelorus Turga, Russia, Early Cretaceous (Aptian)
†Korehelorus Rosse-Guillevic, Rasnitsyn, Nam, & Jouault, 2023 Jinju Formation, South Korea, Early Cretaceous (Albian)
†Laiyanghelorus Zhang 1992 Laiyang Formation, Yixian Formation, China, Early Cretaceous (Aptian)
†Liaoropronia Zhang and Zhang 2001 Yixian Formation, China, Early Cretaceous (Aptian)
†Novhelorus Li et al. 2017 Yixian Formation, China, Early Cretaceous (Aptian)
†Obconohelorus Rasnitsyn 1990 Turga, Russia, Early Cretaceous (Aptian)
†Protocyrtus Karabastau Formation, Kazakhstan, Middle-Late Jurassic (Callovian/Oxfordian) Gurvan-Eren Formation, Mongolia Yixian Formation, China, Turga, Russia, Early Cretaceous (Aptian)
†Sinohelorus Shi et al. 2013 Yixian Formation, China, Early Cretaceous (Aptian)
†Sinuijuhelorus Jon et al. 2019 Sinuiju Formation, North Korea, Early Cretaceous (Aptian)
†Spherogaster Zhang and Zhang 2001 Yixian Formation, China, Early Cretaceous (Aptian)


Izadizadeh, M.; Talebi, A. A.; Van Achterberg, C.; Rakhshani, E. (2015-04-13). "First record of the family Heloridae (Hymenoptera: Proctotrupoidea) from Iran, with description of a new species". Zootaxa. 3946 (4): 577–582. doi:10.11646/zootaxa.3946.4.6. ISSN 1175-5334. PMID 25947710.
"Heloridae Family Information". Retrieved 2018-02-23.
"Heloridae Report". Integrated Taxonomic Information System. Retrieved 2018-02-23.
"Heloridae Overview". Encyclopedia of Life. Retrieved 2018-02-23.

"Browse Heloridae". Catalogue of Life. Retrieved 2018-02-23.

Further reading

Kellogg, Vernon L. (1905). American insects. H. Holt.
Arnett, Ross H. Jr. (2000). American Insects: A Handbook of the Insects of America North of Mexico. Vol. 2nd Edition. CRC Press. ISBN 0-8493-0212-9.
Krombein, Karl V.; Hurd Jr., Paul D. Jr.; Smith, David R.; Burks, B.D. (1979). Catalog of Hymenoptera in America North of Mexico. Smithsonian Institution Press.
Capinera, John L., ed. (2008). Encyclopedia of Entomology. Springer. ISBN 978-1402062421.
Gillott, Cedric (1980). Entomology. Plenum Press. ISBN 0-306-40366-8.
Donald J. Borror; Roger Tory Peterson; Richard E. White (1998). A Field Guide to Insects. Houghton Mifflin.
Goulet H.; Huber J., eds. (1993). "Hymenoptera of the world: an identification guide to families". Agriculture Canada Publication 1894/E.
Sharkey, M.J.; Carpenter, J.M.; Vilhelmsen, L. (2012). "Phylogenetic relationships among superfamilies of Hymenoptera". Cladistics. 28 (1): 80–112. doi:10.1111/j.1096-0031.2011.00366.x. PMID 34861753. S2CID 33628659.

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