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Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Ordo: Coleoptera
Subordo: Polyphaga
Infraordo: Cucujiformia
Superfamilia: Cucujoidea

Familia: Helotidae
Genera (5): Afrohelotina - Helota - Metahelotella - Neohelota - Scrophohelota


Helotidae Reitter & Chapuis, 1876

Kirejtshuk, A.G. 2000: On origin and early evolution of the superfamily Cucujoidea (Coleoptera, Polyphaga). Comments on the family Helotidae. Izvestiya Kharkovskogo Entomologicheskogo Obshchestva, 8(1): 8–38. [in English, with Russian abstract] Abstract
Lawrence, J.F.; Ślipiński, A.; Lee, C.-F. 2010: 10.3. Helotidae Reitter, 1876/Chapuis, 1876. Pp. 292-295 in: Leschen, R.A.B.; Beutel, R.G.; Lawrence, J.F. (volume eds.) Coleoptera, beetles. Volume 2: Morphology and systematics (Elateroidea, Bostrichiformia, Cucujiformia partim). In: Kristensen, N.P. & Beutel, R.G. (eds.) Handbook of zoology. A natural history of the phyla of the animal kingdom. Volume IV. Arthropoda: Insecta. Part 38. Berlin, New York: Walter de Gruyter. ISBN 3110190753 ISBN 9783110190755
Lee, C.-F. 2009: Revision of the family Helotidae (Coleoptera: Cucujoidea) III: The Thibetana group and a checklist of Helota MacLeay species. Annals of the Entomological Society of America, 102: 48–59.
Lee, C.-F. 2010: Revision of the family Helotidae (Coleoptera: Cucujoidea): V. Species group classification of the genus Neohelota Ohta and revisions of N. laevigata and N. helleri species groups. Annals of the Entomological Society of America, 103: 500–510. DOI: 10.1603/AN10003
Lee, C.-f.; Hisamatsu, S.; Yang, P.-s. 2007: Morphology and ontogeny of immature stages of Helotidae based on descriptions of Helota thoracica Ritsema and H. gemmata Gorham (Insecta: Coleoptera: Cucujoidea). Zoological studies, 46: 760–769. PDF
Lee, C.-f.; Satô, M. 2006: The Helotidae of Taiwan (Coleoptera: Cucujoidea). Zoological studies, 45: 529–552. PDF
Lee, C.-F. & P. Votruba, 2011: Revision of Family Helotidae (Coleoptera: Cucujoidea): VI. Candezei Group of the Genus: Neohelota. Annals of the Entomological Society of America 104 (4): 658–665. Abstract: DOI: 10.1603/AN11021.
I.Löbl & A.Smetana (eds). 2007 Catalogue of Palearctic Coleoptera. Vol. 4: Elateroidea, Derodontoidea, Bostrichoidea, Lymexyloidea, Cleroidea and Cucujoidea. Apollo Books, Stenstrup, Denmark ISBN 87-88757-67-6, p. 495
Wegrzynowicz, P. 2000: Catalogue of the Helotidae (Coleoptera: Cucujoidea). Zoologische Mededelingen (Leiden), 73: 391–411. Abstract PDF


Tree of Life Web Project. 2008. Helotidae. Helota. Version 23 June 2008 (temporary) in The Tree of Life Web Project
Atlas of the family Helotidae of Russia

Helotidae is a family of beetles, in the suborder Polyphaga. The family includes about five extant genera, Helota MacLeay, Neohelota Ohta, Afrohelotina Kirejtshuk, Metahelotella Kirejtshuk, and Strophohelota Kirejtshuk.[1][2] Helotidae are found mainly in the Old World tropics and are absent from Australia and Madagascar. The antennae are clubbed on the final three segments and is retractable within grooves under the head. The wings have reduced venation with just 4 anal veins.[3]

A few fossil genera have been described including Palaeohelota from the Lower Cretaceous (Aptian) Yixian Formation in China, Burmahelota, Metahelotella and Trihelota from the Upper Cretaceous (Cenomanian) Burmese amber of Northern Myanmar. Another fossil genus Laodiscis placed in this family originally is now considered of uncertain taxonomic position.[4]


Lee, Chi-Feng; Votruba, Petr (2013). "A revision of the family Helotidae (Coleoptera: Cucujoidea): VIII. The guerinii species-group of the genus Neohelota". Entomologica Basiliensia et Collectionis Frey. 34: 269–308.
Lee, Chi-Feng (2007). "Revision of Family Helotidae (Coleoptera: Cucujoidea):I. Gemmata Group of Genus: Helota". Ann. Entomol. Soc. Am. 100 (5): 623–639.
Crowson, R.A. (1967). The natural classification of the families of Coleoptera. Middlesex: E.W. Classey Ltd. p. 104.

Liu, Zhenhua; Ślipiński, Adam; Ren, Dong; Pang, Hong (2019). "The first Mesozoic Helotidae (Coleoptera: Cucujoidea)". Cretaceous Research. 96: 113–119. doi:10.1016/j.cretres.2018.12.005.

Additional sources

Ritsema C. (1915) A systematic catalogue of the Coleopterous family Helotidae in the Leiden Museum Zoologische Mededelingen, Vol. 1 P. 125-139 PDF
Wegrzynowicz P. (2000) Catalogue of the Helotidae (Coleoptera: Cucujoidea), Zoologische Mededelingen, Vol. 73 P. 391-411 PDF

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