Superregnum: Eukaryota
Cladus: Orthokaryota
Cladus: Neokaryota
Cladus: Scotokaryota
Cladus: Podiata
Cladus: Amorphea
Cladus: Obazoa
Cladus: Opisthokonta
Cladus: Holozoa
Cladus: Filozoa
Cladus: Choanozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: ParaHoxozoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Cladus: Olfactores
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Cladus: Sarcopterygii
Cladus: Rhipidistia
Cladus: Tetrapodomorpha
Cladus: Eotetrapodiformes
Cladus: Elpistostegalia
Superclassis: Tetrapoda
Cladus: Batrachomorpha
Classis: Amphibia
Subclassis: Lissamphibia
Superordo: Batrachia
Cladus: Salientia
Ordo: Anura
Familia: Bufonidae
Genus: Ingerophrynus
Species (12): I. biporcatus – I. celebensis – I. claviger – I. divergens – I. galeatus – I. gollum – I. kumquat – I. ledongensis – I. macrotis – I. parvus – I. philippinicus – I. quadriporcatus
Name
Ingerophrynus Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green & Wheeler, 2006: 219
Type species: Bufo biporcatus Gravenhorst, 1829, by original designation.
References
Primary references
Frost, D.R., Grant, T., Faivovich, J., Bain, R.H., Haas, A., Haddad, C.F.B., de Sá, R.O., Channing, A., Wilkinson, M., Donnellan, S.C., Raxworthy, C.J., Campbell, J.A., Blotto, B.L., Moler, P.E., Drewes, R.C., Nussbaum, R.A., Lynch, J.D., Green, D.M. & Wheeler, W.C. 2006. The amphibian tree of life. Bulletin of the American Museum of Natural History 297: 1–370. DOI: 10.1206/0003-0090(2006)297[0001:TATOL]2.0.CO;2 Paywall; hdl: 2246/5781 Open access. Reference page.
Ingerophrynus is a genus of true toads with 12 species.[2][3] The genus is found in southern Yunnan and Southeast Asia; from Myanmar and Indochina to peninsular Thailand and Malaya, Sumatra, Borneo, Java, Nias Island, Sulawesi, and the Philippines.[2] This genus was established after a major taxonomical revision of frogs in 2006.[1]
Taxonomy and systematics
Ten of the species in this genus were formerly considered species of the genus Bufo, most of them in what had been known as the Bufo biporcatus group; the remaining ones were assigned to this genus based on molecular data.[1] In 2007 a new species, Ingerophrynus gollum, was added to this genus.[4] The sister taxon of Ingerophrynus is Sabahphrynus.[2]
Etymology
The generic name Ingerophrynus honors Robert F. Inger, an American zoologist from the Field Museum of Natural History.[1][5]
Description
The diagnostic characters of the Bufo biporcatus group are the presence of supraorbital, parietal, and supratympanic crests, lack of a tarsal ridge, presence of vocal sacs but absence of melanophores in the surrounding muscle tissue, lack of tibial glands, lack supinator manus humeralis and adductor longus muscles, presence of paired crests on the vertebral column, rugose skull, squamosal bones with broad dorsal otic plates, and smooth palatine bones.[4]
Species
There are 12 species:[2][3]
Binomial name and binomial authority common name
Ingerophrynus biporcatus (Gravenhorst, 1829) crested toad
Ingerophrynus celebensis (Günther, 1859) Sulawesian toad
Ingerophrynus claviger (W. Peters, 1863) Benkulen toad
Ingerophrynus divergens (W. Peters, 1871) Malayan dwarf toad
Ingerophrynus galeatus (Günther, 1864) bony-headed toad
Ingerophrynus gollum L. Grismer, 2007
Ingerophrynus kumquat (Das & Lim, 2001)
Ingerophrynus ledongensis (Fei, Ye & Huang, 2009)
Ingerophrynus macrotis (Boulenger, 1887) big-eared toad
Ingerophrynus parvus (Boulenger, 1887) lesser Malacca toad
Ingerophrynus philippinicus (Boulenger, 1887) Philippine toad
Ingerophrynus quadriporcatus (Boulenger, 1887) greater Malacca toad
Nota bene: A binomial authority in parentheses indicates that the species was originally described in a genus other than Ingerophrynus.
References
Frost, D.R.; Grant, T.; Faivovich, J.; Bain, R.H.; Haas, A.; Haddad, C.L.F.B.; Sá, R.O.; Channing, A.; Wilkinson, M.; Donnellan, S.C.; Raxworthy, C.J.; Campbell, J.A.; Blotto, B.L.; Moler, P.; Drewes, R.C.; Nussbaum, R.A.; Lynch, J.D.; Green, D.M.; Wheeler, W.C. (2006). "The amphibian tree of life". Bulletin of the American Museum of Natural History. 297: 1–291. doi:10.1206/0003-0090(2006)297[0001:TATOL]2.0.CO;2. hdl:2246/5781.
Frost, Darrel R. (2018). "Ingerophrynus Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006". Amphibian Species of the World: an Online Reference. Version 6.0. American Museum of Natural History. Retrieved 7 March 2018.
"Bufonidae". AmphibiaWeb. University of California, Berkeley. 2018. Retrieved 7 March 2018.
Grismer, L. Lee (2007). "A new species of Ingerophrynus (Anura: Bufonidae) from a lowland rain forest in southern peninsular Malaysia". Journal of Herpetology. 41 (2): 225–230. doi:10.1670/0022-1511(2007)41[225:ansoia]2.0.co;2. JSTOR 4498578. S2CID 198157730.
Beolens, Bo; Watkins, Michael; Grayson, Michael (2013). The Eponym Dictionary of Amphibians. Exeter, England: Pelagic Publishing. pp. 101–102. ISBN 978-1-907807-42-8. (ePub).
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