Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: ParaHoxozoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Cladus: Olfactores
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Cladus: Sarcopterygii
Cladus: Rhipidistia
Cladus: Tetrapodomorpha
Cladus: Eotetrapodiformes
Cladus: Elpistostegalia
Superclassis: Tetrapoda
Cladus: Batrachomorpha
Classis: Amphibia
Subclassis: Lissamphibia
Superordo: Batrachia
Cladus: Salientia
Ordo: Anura
Familia: Megophryidae
Subfamilia: Leptobrachiinae
Genus: Leptobrachella
Species (107): L. aerea – L. alpina – L. applebyi – L. arayai – L. ardens – L. aspera – L. aurantirosea – L. baluensis – L. bashaensis – L. bidoupensis – L. bijie – L. bondangensis – L. botsfordi – L. bourreti – L. brevicrus – L. chishuiensis – L. crocea – L. damingshanensis – L. dayaoshanensis – L. dong – L. dorsospina – L. dringi – L. dushanensis – L. eos – L. feii – L. firthi – L. flaviglandulosa – L. fritinniens – L. fuliginosa – L. fusca – L. gracilis – L. graminicola – L. guinanensis – L. hamidi – L. heteropus – L. huynhi – L. isos – L. itiokai – L. jinshaensis – L. jinyunensis – L. juliandringi – L. kajangensis – L. kalonensis – L. kecil – L. khasiorum – L. korifi – L. lateralis – L. laui – L. liui – L. macrops – L. maculosa – L. mangshanensis – L. maoershanensis – L. marmorata – L. maura – L. melanoleuca – L. melica – L. minima – L. mjobergi – L. murphyi – L. nahangensis – L. namdongensis – L. natunae – L. neangi – L. niveimontis – L. nokrekensis – L. nyx – L. oshanensis – L. pallida – L. palmata – L. parva – L. pelodytoides – L. petrops – L. phiadenensis – L. phiaoacensis – L. picta – L. pingbianensis – L. platycephala – L. pluvialis – L. puhoatensis – L. purpuraventra – L. purpurus – L. pyrrhops – L. rowleyae – L. sabahmontana – L. serasanae – L. shangsiensis – L. shimentaina – L. shiwandashanensis – L. sinorensis – L. sola – L. suiyangensis – L. sungi – L. tadungensis – L. tamdil – L. tengchongensis – L. tuberosa – L. ventripunctata – L. verrucosa – L. wuhuangmontis – L. wulingensis – L. wumingensis – L. yeae – L. yingjiangensis – L. yunkaiensis – L. yunyangensis – L. zhangyapingi
Name
Leptobrachella Smith, 1925: 27
Type species: Leptobrachella mjobergi Smith, 1925, by monotypy.
Synonyms
Nesobia Van Kampen, 1923: 6 [nomen praeoccupatum pro Nesobia Ancey, 1887]
Type species: Leptobrachium natunae Günther, 1895, by monotypy.
Leptolalax Dubois, 1980: 476 [as a subgenus of Leptobrachium; elevate to genus by Dubois (1983: 147); synonymised by Chen et al. (2018: 162)]
Type species: Leptobrachium gracile Günther, 1872, by original designation.
Lalax Delorme, Dubois, Grosjean & Ohler, 2006: 14 [as a subgenus of Leptolalax; nomen praeoccupatum pro †Lalax Hamilton, 1990]
Type species: Leptobrachium bourreti Dubois, 1983, by original designation.
Lalos Dubois, Grosjean, Ohler, Adler & Zhao, 2010: 66 [nomen substitutum pro Lalax Delorme, Dubois, Grosjean & Ohler, 2006]
Type species: Leptobrachium bourreti Dubois, 1983, by original designation.
References
Primary references
Smith, M.A. 1925. Contributions to the herpetology of Borneo. Sarawak Museum Journal 3: 15–34.
van Kampen, P.N. 1923. The Amphibia of the Indo-Australian Archipelago. E.J. Brill Ltd: Leiden. 304 pp. BHL Reference page.
Dubois, A. 1980. Notes sur la systematique et la repartition des amphibiens anoures de Chine et des regions avoisinantes IV. Classification generique et subgenerique des Pelobatidae Megophryinae. Bulletin Mensuel de la Société Linnéenne de Lyon 49: 469–482.
Dubois, A. 1983. Note préliminaire sur le genre Leptolalax Dubois, 1980 (Amphibiens, Anoures), avec diagnose d'une espèce nouvelle du Vietnam. Alytes 2: 147–153.
Delorme, M., Dubois, A., Grosjean, S. & Ohler, A. 2006. Une nouvelle ergotaxinomie des Megophryidae (Amphibia, Anura). Alytes 24(1-4): 6–21. Reference page.
Dubois, A., Grosjean, S., Ohler, A., Adler, K. & Zhao, E.-M. 2010. The nomenclatural status of some generic nomina of Megophryidae (Amphibia: Anura). Zootaxa 2493(1): 66–68. DOI: 10.11646/zootaxa.2493.1.6 Paywall Reference page.
Chen, J., Poyarkov Jr., N.A., Suwannapoom, C., Lathrop, A., Wu, Y.-H., Zhou, W.-W., Yuan, Z.-Y., Jin, J.-Q., Chen, H.-M., Liu, H.-Q., Nguyen, T.Q., Nguyen, S.N., Duong, T.V., Eto, K., Nishikawa, K., Matsui, M., Orlov, N.L., Stuart, B.L., Brown, R.M., Rowley, J.J.L., Murphy, R.W., Wang, Y.-Y. & Che, J. 2018. Large-scale phylogenetic analyses provide insights into unrecognized diversity and historical biogeography of Asian leaf-litter frogs, genus Leptolalax (Anura: Megophryidae). Molecular Phylogenetics and Evolution 124: 162–171. DOI: 10.1016/j.ympev.2018.02.020 Paywall Reference page.
Additional references
Duong, T.V., Do, D.T., Ngo, C.D., Nguyen, T.Q. & Poyarkov, Jr., N.A. 2018. A new species of the genus Leptolalax (Anura: Megophryidae) from southern Vietnam. Zoological Research 39(3): 181–196. DOI: 10.24272/j.issn.2095-8137.2018.009 Open access Reference page.
Eto, K., Matsui, M., Hamidy, A., Munir, M. & Iskandar, D. 2018. Two New Species of the Genus Leptobrachella (Amphibia: Anura: Megophryidae) from Kalimantan, Indonesia. Current Herpetology 37(2): 95–105. DOI: 10.5358/hsj.37.95 Paywall Reference page.
Hou, Y-M., Zhang, M-F., Hu, F., Li, S-Y., Shi, S-C., Chen, J., Mo, X-Y. & Wang, B. 2018. A new species of the genus Leptolalax (Anura, Megophryidae) from Hunan, China. Zootaxa 4444(3): 247–266. DOI: 10.11646/zootaxa.4444.3.2 Paywall Reference page.
Nguyen, L.T., Poyarkov, N.A. Jr., Le, D.T., Vo, B.D., Phan, H.T., Duong, T.V., Murphy, R.W. & Nguyen, S.N. 2018. A new species of Leptolalax (Anura: Megophryidae) from Son Tra Peninsula, central Vietnam. Zootaxa 4388(1): 1–21. DOI: 10.11646/zootaxa.4388.1.1 Paywall Reference page.
Yang, J.-H., Zeng, Z.-C. & Wang, Y.-Y. 2018. Description of two new sympatric species of the genus Leptolalax (Anura: Megophryidae) from western Yunnan of China. PeerJ 6: e4586. DOI: 10.7717/peerj.4586 Open access Reference page.
Wang, J., Yang, J-H., Li, Y., Lyu, Z-T., Zeng, Z-C., Liu, Z-Y., Ye, Y-H. & Wang, Y-Y. 2018. Morphology and molecular genetics reveal two new Leptobrachella species in southern China (Anura, Megophryidae). ZooKeys 776: 105–137. DOI: 10.3897/zookeys.776.22925 Open access Reference page.
Chen, W-C., Liao, X-W., Zhou, S-C. & Mo, Y-M. 2019. A new species of Leptobrachella (Anura: Megophryidae) from southern Guangxi, China. Zootaxa 4563(1): 67–82. DOI: 10.11646/zootaxa.4563.1.3 Paywall Reference page.
Hoang, V.C., Nguyen, T.T., Luu, Q.V., Nguyen, Q.T. & Jiang, J.-P. 2019. A new species of Leptobrachella Smith 1925 (Anura: Megophryidae) from Thanh Hoa Province, Vietnam. The Raffles Bulletin of Zoology 67: 536–556. DOI: 10.26107/RBZ-2019-0042 Open access Reference page.
Wang, J., Li, Y-L., Li, Y., Chen, H-H., Zeng, Y-J., Shen, J-M. & Wang, Y-Y. 2019. Morphology, molecular genetics, and acoustics reveal two new species of the genus Leptobrachella from northwestern Guizhou Province, China (Anura, Megophryidae). Zookeys 848: 119–154. DOI: 10.3897/zookeys.848.29181 Open access Reference page.
Chen, J.-M., Xu, K., Poyarkov, Jr., N.A., Wang, K., Yuan, Z.-H., Hou, M., Suwannapoom, C., Wang, J. & Che, J. 2020. How little is known about “the little brown frogs”: description of three new species of the genus Leptobrachella (Anura: Megophryidae) from Yunnan Province, China. Zoological Research 41(3): 292–313. DOI: 10.24272/j.issn.2095-8137.2020.036 Open access Reference page.
Li, S-Z., Liu, J., Wei, G. & Wang, B. 2020. A new species of the Asian leaf litter toad genus Leptobrachella (Amphibia, Anura, Megophryidae) from southwest China. ZooKeys 943: 91–118. DOI: 10.3897/zookeys.943.51572 Open access Reference page.
Luo, T., Xiao, N., Gao, K. & Zhou, J. 2020. A new species of Leptobrachella (Anura, Megophryidae) from Guizhou Province, China. ZooKeys 923: 115–140. DOI: 10.3897/zookeys.923.47172 Open access Reference page.
Lyu, J-C., Dai, L-L., Wei, P-F., He, Y-H., Yuan, Z-Y., Shi, W-L., Zhou, S-L., Ran, S-Y., Kuang, Z-F., Guo, X., Wei, G. & Yuan, G. 2020. A new species of the genus Leptobrachella Smith, 1925 (Anura, Megophryidae) from Guizhou, China. ZooKeys 1008: 139–157. DOI: 10.3897/zookeys.1008.56412 Open access Reference page.
Qian, T.-Y., Xia, X., Cao, Y., Xiao, N.-W. & Yang, D.-D. 2020. A new species of Leptobrachella (Anura: Megophryidae) Smith, 1925 from Wuling Mountains in Hunan Province, China. Zootaxa 4816(4): 491–526. DOI: 10.11646/zootaxa.4816.4.4 Paywall Reference page.
Stuart, B.L. & Rowley, J.J.L. 2020. A new Leptobrachella (Anura: Megophryidae) from the Cardamom Mountains of Cambodia. Zootaxa 4834(4): 556–572. DOI: 10.11646/zootaxa.4834.4.4 Paywall Reference page.
Wang, J., Lyu, Z-T., Qi, S., Zeng, Z-C., Zhang, W-X., Lu, L-S. & Wang, Y-Y. 2020. Two new Leptobrachella species (Anura, Megophryidae) from the Yunnan-Guizhou Plateau, southwestern China. ZooKeys 995: 97–125. DOI: 10.3897/zookeys.995.55939 Open access Reference page.
Chen, J.-M., Suwannapoom, C., Wu, Y.-H., Poyarkov, Jr., N.A., Xu, K., Pawangkhanant, P. & Che, J. 2021. Integrative taxonomy reveals a new species of Leptobrachella (Anura: Megophryidae) from the mountains of northern Thailand. Zootaxa 5052(2): 191–214. DOI: 10.11646/zootaxa.5052.2.2 Paywall Reference page.
Chen, W.-C., Peng, W.-X., Pan, W.-H., Liao, N.-Y., Liu, Y.-J. & Huang, Y. 2021. A new species of Leptobrachella Smith 1925 (Anura: Megophryidae) from southern Guangxi, China. Zootaxa 5020(3): 581–596. DOI: 10.11646/zootaxa.5020.3.8 Paywall Reference page.
Chen, W.-C., Yu, G.-D., Cheng, Z.-Y., Meng, T., Wei, H., Zhou, G.-Y. & Lu, Y.-W. 2021. A new species of Leptobrachella (Anura: Megophryidae) from central Guangxi, China. Zoological Research 42(6): 783–788. DOI: 10.24272/j.issn.2095-8137.2021.179 Open access Reference page.
Cheng, Y.-L., Shi, S.-C., Li, J., Liu, J., Li, S.-Z. & Wang, B. 2021. A new species of the Asian leaf litter toad genus Leptobrachella Smith, 1925 (Anura, Megophryidae) from northwest Guizhou Province, China. ZooKeys 1021: 81–107. DOI: 10.3897/zookeys.1021.60729 Open access Reference page.
Nguyen, L.T., Tapley, B., Nguyen, C.T., Luong, H.V. & Rowley, J.J.L. 2021. A new species of Leptobrachella (Anura, Megophryidae) from Mount Pu Ta Leng, northwest Vietnam. Zootaxa 3(5016): 301–332. DOI: 10.11646/zootaxa.3.5016.1 Paywall Reference page.
Shi, S.-C., Hou, Y.-M., Song, Z.-B., Jiang, J.-P. & Wang, B. 2021. A new Leaf Litter Toad of Leptobrachella Smith, 1925 (Anura, Megophryidae) from Sichuan Province, China with supplementary description of L. oshanensis. Asian Herpetological Research 12(2): 143–166. DOI: 10.16373/j.cnki.ahr.200118 Open access Reference page.
Lin, S.-S., Li, Y-H., Lu, Y.-H., Su, H-L., Wu, S.-B., Zhang, Q.-Q., Mo, M.-J., Xiao, S.-J., Pan, Z., Pan, H.-J., Zeng, Z-C. & Wang, J. 2022. A new species of the genus Leptobrachella (Anura, Megophryidae) from northwestern Guangdong Province, China. Herpetozoa 35: 165–178. DOI: 10.3897/herpetozoa.35.e89981 Open access Reference page.
Luo, T., Wang, W.-F., Peng, D., Lei, B., Deng, H.-Q., Ji, S.-N., Huang, H.-Q. & Zhou, J. 2022. A new species of the Asian Leaf Litter Toad genus Leptobrachella (Amphibia, Anura, Megophryidae) from Chongqing City, Southwest China. Asian Herpetological Research 13(2): 75–95. DOI: 10.16373/j.cnki.ahr.210052 Open access Reference page.
Wang, J., Qi, S., Dai, K.-Y., Lyu, Z.-T., Zeng, Z.-C., Chen, H.-H., Li, Y.-Q., Zhao, Y.-Y., Wang, Y.-Z. & Wang, Y.-Y. 2022. A new Leptobrachella species (Anura, Megophryidae) from South China, with comments on the taxonomic status of L. chishuiensis and L. purpurus. Zoosystematics and Evolution 98: 165–180. DOI: 10.3897/zse.98.73162 Open access Reference page.
Matsui, M., Panha, S. & Eto, K. 2023. Two new species of Leptobrachella from northern Thailand (Amphibia, Anura, Megophryidae). Current Herpetology 42(1): 83–97. DOI: 10.5358/hsj.42.83 Open access Reference page.
Chen, W-C., Peng, W-X., Li, P. & Yu, G-D. 2023. A new species of the genus Leptobrachella Smith 1925 (Amphibia, Anura, Megophryidae) from Guangxi, China. Zookeys 1178: 1–16. DOI: 10.3897/zookeys.1178.106038 Open access Reference page.
Liu, J., Shi, S-C., Li, S., Zhang, M-F., Xiang, S-J., Wei, G. & Wang, B. 2023. A new Asian leaf litter toad of the genus Leptobrachella (Amphibia, Anura, Megophryidae) from central south China. Zookeys 1149: 103–134. DOI: 10.3897/zookeys.1149.85895 Open access Reference page.
Chen, W-C., Li, P., Peng, W-X., Liu, Y-J. & Huang, Y. 2024. The fourth species of Leptobrachella (Anura, Megophryidae) found at Shiwandashan National Nature Reserve, Guangxi, China. ZooKeys 1192: 257–279. DOI: 10.3897/zookeys.1192.98352 Open access Reference page.
Yu, G-D., Qin, K., Meng, T., Li, P., Peng, W-X. & Chen, W-C. 2024. A new species of the genus Leptobrachella (Amphibia, Anura, Megophryidae) from Dayaoshan National Nature Reserve, Guangxi, China. ZooKeys 1219: 105–122. DOI: 10.3897/zookeys.1219.121027 Open access Reference page.
Vernacular names
English: Asian Leaf Litter Toads
Leptobrachella is a genus of frogs in the family Megophryidae. Members of Leptobrachella are found throughout Asia including on Borneo and the Natuna Islands. They are sometimes referred to as Borneo frogs, slender-armed frogs,[1] or dwarf litter frogs.[2][3] The genus contains over 82 species with 25 found in China alone.
Description
Leptobrachella are small frogs that are not easily seen as they are well camouflaged on the ground. However, their advertisement call is loud, and they can be abundant along streams.[3]
The tadpoles of Leptobrachella are unusual in their vermiform or eel-like appearance. The transition from the narrow, cylindrical trunk into the strong tail is nearly seamless, and the tail fin is very low. This body shape is interpreted as an adaptation to a fossorial life style: Leptobrachella tadpoles live in the gravel beds of small streams. In Leptobrachella mjobergi where more detailed observations have been made, tadpoles have unusually mobile head and trunk. While smaller tadpoles seem to use existing interstitial spaces, larger ones can actively push their way through gravel.[4]
Diet and reproduction
Leptobrachella are carnivorous amphibians, consuming about any insect, such as mosquitoes, spiders, grasshoppers, and butterflies (Cheng, 2021). In terms of reproduction, they lay their eggs in the water, such as in ponds. Eventually those eggs hatch into larvae called tadpoles. These tadpoles have tails andinternal gills (Cheng, 2021).
Distribution
Leptobrachella are usually found in the forests of Southeast Asia; however they can be found in southwestern Cambodia and adjacent to Thailand (Cheng, 2021).
Species
The following 108 species are recognised in the genus Leptobrachella:[1]
Leptobrachella aerea (Rowley et al., 2010)
Leptobrachella alpina (Fei, Ye, and Li, 1990)
Leptobrachella applebyi (Rowley and Cao, 2009)
Leptobrachella arayai (Matsui, 1997)
Leptobrachella ardens (Rowley et. al, 2016)
Leptobrachella aspera Wang, Lyu, Qi, and Wang, 2020
Leptobrachella aurantirosea Ninh et. al, 2024 )
Leptobrachella baluensis Smith, 1931
Leptobrachella bashaensis Lyuet. al, 2020[5]
Leptobrachella bidoupensis (Rowley, Le, Tran, and Hoang, 2011)
Leptobrachella bijie Wang, Li, Li, Chen, and Wang, 2019
Leptobrachella bondangensis Eto et al., 2018
Leptobrachella botsfordi (Rowley, Dau, and Nguyen, 2013)
Leptobrachella bourreti (Dubois, 1983)
Leptobrachella brevicrus Dring, 1984
Leptobrachella chishuiensis Li, Liu, Wei, and Wang, 2020
Leptobrachella crocea (Rowley, Hoang, Le, Dau, and Cao, 2010)
Leptobrachella damingshanensis Chen et al., 2021
Leptobrachella dayaoshanensis Chen, Yu, Meng, and Qin, 2024
Leptobrachella dong Liu, Shi, Li, Zhang, Xiang, Wei, and Wang, 2023
Leptobrachella dorsospina Wang, Lyu, Qi, and Wang, 2020
Leptobrachella dringi (Dubois, 1987)
Leptobrachella dushanensis Li, Li, Cheng, Liu, Wei, Wang, 2024
Leptobrachella eos (Ohler et. al, 2011)
Leptobrachella feii Chen, Yuan, and Che, 2020
Leptobrachella firthi (Rowley, Hoang, Dau, Le, and Cao, 2012)
Leptobrachella flaviglandulosa Chen, Wang, and Che, 2020
Leptobrachella fritinniens (Dehling and Matsui, 2013)
Leptobrachella fuliginosa (Matsui, 2006)
Leptobrachella fusca Eto, Matsui, Hamidy, Munir, and Iskandar, 2018
Leptobrachella gracilis (Günther, 1872)
Leptobrachella graminicola Nguyen et al., 2021
Leptobrachella guinanensis Chen, Li, Peng, and Liu, 2024
Leptobrachella hamidi (Matsui, 1997)
Leptobrachella heteropus (Boulenger, 1900)
Leptobrachella huynhi Hoang et. al, 2024
Leptobrachella isos (Rowley et. al, 2015)
Leptobrachella itiokai Eto, Matsui, and Nishikawa, 2016
Leptobrachella jinshaensis Cheng et al., 2021
Leptobrachella jinyunensis Shi, Shen, Wang, Jiang, and Wang, 2023
Leptobrachella juliandringi Eto, Matsui, and Nishikawa, 2015
Leptobrachella kajangensis Grismer, Grismer, and Youmans, 2004
Leptobrachella kalonensis Rowley et al., 2016
Leptobrachella kecil (Matsui, Belabut, Ahmad, and Yong, 2009)
Leptobrachella khasiorum (Das, Tron, Rangad, and Hooroo, 2010
Leptobrachella korifi Matsui, Panha, and Eto, 2023
Leptobrachella lateralis (Anderson, 1871)
Leptobrachella laui (Sung, Yang, and Wang, 2014)
Leptobrachella liui Fei and Ye, 1990
Leptobrachella macrops (Duong, Do, Ngo, Nguyen, and Poyarkov, 2018)
Leptobrachella maculosa (Rowley et al., 2016)
Leptobrachella mangshanensis (Hou et al., 2018)
Leptobrachella maoershanensis (Yuan, Sun, Chen, Rowley, and Che, 2017)
Leptobrachella marmorata (Matsui, Zainudin, and Nishikawa, 2014)
Leptobrachella maura (Inger, Lakim, Biun, and Yambun, 1997)
Leptobrachella melanoleuca (Matsui, 2006)
Leptobrachella melica (Rowley, Stuart, Neang, and Emmett, 2010)
Leptobrachella minima Taylor, 1962
Leptobrachella mjobergi Smith, 1925
Leptobrachella murphyi Chen et al., 2021
Leptobrachella nahangensis (Lathrop, Murphy, Orlov, and Ho, 1998)
Leptobrachella namdongensis Hoang et al., 2019
Leptobrachella natunae (Günther, 1895)
Leptobrachella neangi Stuart and Rowley, 2020
Leptobrachella niveimontis Chen, Poyarkov, Yuan, and Che, 2020
Leptobrachella nokrekensis Mathew and Sen, 2010
Leptobrachella nyx Ohler et al., 2011
Leptobrachella oshanensis Liu, 1950
Leptobrachella pallida (Rowley et al., 2016)
Leptobrachella palmata Inger and Stuebing, 1992
Leptobrachella parva Dring, 1984
Leptobrachella pelodytoides (Boulenger, 1893)
Leptobrachella petrops (Rowley et al., 2017)
Leptobrachella phiadenensis Luong et al., 2023
Leptobrachella phiaoacensis Luong et al., 2023
Leptobrachella picta Malkmus, 1992
Leptobrachella pingbianensis (Rao, Hui, Zhu, and Ma, 2022 "2020")
Leptobrachella platycephala (Dehling, 2012)
Leptobrachella pluvialis (Ohler, Marquis, Swan, and Grosjean, 2000)
Leptobrachella puhoatensis (Rowley, Dau, and Cao, 2017)
Leptobrachella purpuraventra Wang, Li, Li, Chen, and Wang, 2019
Leptobrachella purpuris (Yang, Zeng, and Wang, 2018)
Leptobrachella pyrrhops (Poyarkov et al., 2015)
Leptobrachella rowleyae (Nguyen et al., 2018)
Leptobrachella sabahmontana (Matsui, Nishikawa, and Yambun, 2014)
Leptobrachella serasanae Dring, 1984
Leptobrachella shangsiensis Chen, Liao, Zhou, and Mo, 2019
Leptobrachella shimentaina Wang, Lyu, and Wang, 2022
Leptobrachella shiwandashanensis Chen et al., 2021
Leptobrachella sinorensis Matsui, Panha, and Eto, 2023
Leptobrachella sola (Matsui, 2006)
Leptobrachella suiyangensis Luo, Xiao, Gao, and Zhou, 2020
Leptobrachella sungi (Lathrop, Murphy, Orlov, and Ho, 1998)
Leptobrachella tadungensis (Rowley et al., 2016)
Leptobrachella tamdil (Sengupta, Sailo, Lalremsanga, Das, and Das, 2010
Leptobrachella tengchongensis (Yang, Wang, Chen, and Rao, 2016)
Leptobrachella tuberosa (Inger, Orlov, and Darevsky, 1999)
Leptobrachella ventripunctata (Fei, Ye, and Li, 1990)
Leptobrachella verrucosa Wang, Zeng, Lin, and Li, 2022
Leptobrachella wuhuangmontis Wang, Yang, and Wang, 2018
Leptobrachella wulingensis Qian, Xiao, Cao, Xiao, and Yang, 2020
Leptobrachella wumingensis Chen, Peng, Li, and Yu, 2023
Leptobrachella xishuiensis Luo, Zhao, Wang, Lan, Xiao, Deng, Xiao, and Zhou, 2025
Leptobrachella yeae Shi, Hou, Song, Jiang, and Wang, 2021
Leptobrachella yingjiangensis (Yang, Zeng, and Wang, 2018)
Leptobrachella yunkaiensis Wang, Li, Lyu, and Wang, 2018
Leptobrachella yunyangensis Luo, Deng, and Zhou, 2022
Leptobrachella zhangyapingi (Jiang et al., 2013)
References
Cheng, Shi, S.-C., Li, J., Liu, J., Li, S.-Z., & Wang, B. (2021). A new species of the Asian leaf litter toad genus Leptobrachella Smith, 1925 (Anura, Megophryidae) from northwest Guizhou Province, China. ZooKeys, 1021(8), 81–107. https://doi.org/10.3897/zookeys.1021.60729
Frost, Darrel R. (2024). "Leptobrachella Smith, 1925". Amphibian Species of the World: an Online Reference. Version 6.2. American Museum of Natural History. Retrieved 2 January 2025.
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