Accipitrinae

Life-forms

Superregnum: Eukaryota
Cladus: Orthokaryota
Cladus: Neokaryota
Cladus: Scotokaryota
Cladus: Podiata
Cladus: Amorphea
Cladus: Obazoa
Cladus: Opisthokonta
Cladus: Holozoa
Cladus: Filozoa
Cladus: Choanozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: ParaHoxozoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Cladus: Olfactores
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Cladus: Sarcopterygii
Cladus: Rhipidistia
Cladus: Tetrapodomorpha
Cladus: Eotetrapodiformes
Cladus: Elpistostegalia
Cladus: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Cladus: Sauropsida
Cladus: Reptilia
Cladus: Eureptilia
Cladus: Romeriida
Subclassis: Diapsida
Cladus: Neodiapsida
Cladus: Sauria
Cladus: Archelosauria
Cladus: Archosauromorpha
Cladus: Crocopoda
Cladus: Archosauriformes
Cladus: Eucrocopoda
Cladus: Archosauria
Cladus: Avemetatarsalia
Cladus: Ornithodira
Cladus: Dinosauromorpha
Cladus: Dinosauriformes
Cladus: Dracohors
Cladus: Dinosauria
Cladus: Saurischia
Cladus: Theropoda
Cladus: Neotheropoda
Cladus: Averostra
Cladus: Tetanurae
Cladus: Avetheropoda
Cladus: Coelurosauria
Cladus: Tyrannoraptora
Cladus: Maniraptoromorpha
Cladus: Maniraptoriformes
Cladus: Maniraptora
Cladus: Pennaraptora
Cladus: Paraves
Cladus: Eumaniraptora
Cladus: Avialae
Subclassis: Aves
Cladus: Euavialae
Cladus: Avebrevicauda
Cladus: Pygostylia
Cladus: Ornithothoraces
Cladus: Euornithes
Cladus: Ornithuromorpha
Cladus: Ornithurae
Cladus: Carinatae
Parvclassis: Neornithes
Infraclassis: Neognathae
Cladus: Neoaves
Ordo: Accipitriformes
Familia: Accipitridae
Subfamilia: Accipitrinae
Genera: Accipiter – Aerospiza – Astur – Circus – Erythrotriorchis – Kaupifalco – Megatriorchis – Melierax – Micronisus – Tachyspiza – Urotriorchis
Name

Accipitrinae Vieillot, 1816
References

Vieillot, L.P. 1816. Analyse d'une nouvelle ornithologie élémentaire. 70 pp. Paris: Deterville. p. 22 "Accipitrini" BHL Reference page.

Vernacular names
ak: Akroma
Deutsch: Habicht
Ελληνικά: Γεράκι
English: Hawks
español: Gavilanes
Nederlands: Havik
português: Gavião
русский: Ястребы
svenska: Hökar
ไทย: เหยี่ยว

The Accipitrinae are the subfamily of the Accipitridae often known as the "true" hawks. The subfamily contains 73 species that are divided into 11 genera. It includes the genus Accipiter which formerly included many more species. The large genus was found to be non-monophyletic and was split into several new or resurrected genera. The birds in this subfamily are primarily woodland birds that hunt by sudden dashes from a concealed perch, with long tails, broad wings and high visual acuity facilitating this lifestyle.

Hawks, including the accipitrines, are believed to have vision several times sharper than humans, in part because of the great number of photoreceptor cells in their retinas (up to 1,000,000 per square mm, against 200,000 for humans), a very high number of nerves connecting the receptors to the brain, and an indented fovea, which magnifies the central portion of the visual field.

A series of molecular phylogenetic studies found that the genus Accipiter was non-monophyletic.[1] The results of a densely sampled 2024 study of the Accipitridae allowed the generic boundaries to be redefined.[2] To create monophyletic genera, species were moved from Accipiter to five new or resurrected genera leaving only 9 species in Accipiter.[3] The southeast Asian crested goshawk and the Sulawesi goshawk were found to be only distantly related to other species in Accipiter. They were moved to a resurrected genus Lophospiza, the only genus placed in the new subfamily Lophospizinae. Similarly, the very small south America tiny hawk and semicollared hawk were found to be only distantly related to species in Accipiter. They were moved to a newly erected genus Microspizias which together with Harpagus is placed in the subfamily Harpaginae. The genera Circus, Megatriorchis, and Erythrotriorchis were found to be nested within Accipiter. Rather than subsuming these genera into an expanded Accipiter, species were moved from Accipiter to the resurrected genera Aerospiza, Tachyspiza and Astur.[3]

The genus level cladogram of the Accipitrinae shown below is based on a molecular phylogenetic study of the Accipitridae by Therese Catanach and collaborators that was published in 2024.[2] The number of species in each genus is based on the list maintained by Frank Gill, Pamela C. Rasmussen and David Donsker on behalf of the International Ornithological Committee (IOC).[3]

Species

The subfamily Accipitrinae contains 73 species that are arranged into 11 genera:[3]

Image	Genus	Living Species
	Micronisus G.R. Gray, 1840	Gabar goshawk, Micronisus gabar
	Urotriorchis Sharpe, 1874	Long-tailed hawk, Urotriorchis macrourus
	Melierax G.R. Gray, 1840	Dark chanting goshawk, Melierax metabates Eastern chanting goshawk, Melierax poliopterus Pale chanting goshawk, Melierax canorus
	Kaupifalco Bonaparte, 1854	Lizard buzzard, Kaupifalco monogrammicus
	Aerospiza Roberts, 1922	Red-chested goshawk, Aerospiza toussenelii African goshawk, Aerospiza tachiro Chestnut-flanked sparrowhawk, Aerospiza castanilius
	Tachyspiza Kaup, 1844	Shikra, Tachyspiza badia Nicobar sparrowhawk, Tachyspiza butleri Levant sparrowhawk, Tachyspiza brevipes Chinese sparrowhawk, Tachyspiza soloensis Frances's sparrowhawk, Tachyspiza francesiae Spot-tailed sparrowhawk, Tachyspiza trinotata Grey goshawk, Tachyspiza novaehollandiae Variable goshawk, Tachyspiza hiogaster Brown goshawk, Tachyspiza fasciata Black-mantled goshawk, Tachyspiza melanochlamys Pied goshawk, Tachyspiza albogularis White-bellied goshawk, Tachyspiza haplochroa Fiji goshawk, Tachyspiza rufitorques Moluccan goshawk, Tachyspiza henicogramma Slaty-mantled goshawk, Tachyspiza luteoschistacea Imitator goshawk, Tachyspiza imitator Grey-headed goshawk, Tachyspiza poliocephala New Britain goshawk, Tachyspiza princeps Red-thighed sparrowhawk, Tachyspiza erythropus Little sparrowhawk, Tachyspiza minulla Japanese sparrowhawk, Tachyspiza gularis Besra, Tachyspiza virgata Dwarf sparrowhawk, Tachyspiza nanus Rufous-necked sparrowhawk, Tachyspiza erythrauchen Collared sparrowhawk, Tachyspiza cirrocephala New Britain sparrowhawk, Tachyspiza brachyura Vinous-breasted sparrowhawk, Tachyspiza rhodogaster
	Erythrotriorchis Sharpe, 1875	Chestnut-shouldered goshawk, Erythrotriorchis buergersi Red goshawk, Erythrotriorchis radiatus
	Accipiter Brisson, 1760	Madagascar sparrowhawk, Accipiter madagascariensis Ovambo sparrowhawk, Accipiter ovampensis Eurasian sparrowhawk, Accipiter nisus Rufous-breasted sparrowhawk, Accipiter rufiventris Grey-bellied hawk, Accipiter poliogaster Sharp-shinned hawk, Accipiter striatus White-breasted hawk, Accipiter chionogaster Plain-breasted hawk, Accipiter ventralis Rufous-thighed hawk, Accipiter erythronemius
	Astur Lacépède, 1799	Cooper's hawk, Astur cooperii Gundlach's hawk, Astur gundlachi Bicolored hawk, Astur bicolor Chilean hawk, Astur chilensis Black sparrowhawk, Astur melanoleucus Henst's goshawk, Astur henstii Eurasian goshawk, Astur gentilis American goshawk, Astur atricapillus Meyer's goshawk, Astur meyerianus
	Megatriorchis Salvadori & D'Albertis, 1875	Doria's hawk, Megatriorchis doriae
	Circus Lacépède, 1799	Western marsh harrier, Circus aeruginosus Eastern marsh harrier, Circus spilonotus Papuan harrier, Circus spilothorax Swamp harrier, Circus approximans African marsh harrier, Circus ranivorus Réunion harrier, Circus maillardi Malagasy harrier, Circus macrosceles Long-winged harrier, Circus buffoni Spotted harrier, Circus assimilis Black harrier, Circus maurus Hen harrier, Circus cyaneus Northern harrier, Circus hudsonius Cinereous harrier, Circus cinereus Pallid harrier, Circus macrourus Pied harrier, Circus melanoleucos Montagu's harrier, Circus pygargus

Hawks and humans

Hawks are sometimes used in falconry, a sport in which trained birds of prey are flown at small game for sport.
References

Mindell, D.; Fuchs, J.; Johnson, J. (2018). "Phylogeny, taxonomy, and geographic diversity of diurnal raptors: Falconiformes, Accipitriformes, and Cathartiformes". In Sarasola, J.H.; Grange, J.M.; Negro, J.J. (eds.). Birds of Prey: Biology and conservation in the XXI century. Cham, Switzerland: Springer. pp. 3–32. ISBN 978-3-319-73744-7.
Catanach, T.A.; Halley, M.R.; Pirro, S. (2024). "Enigmas no longer: using ultraconserved elements to place several unusual hawk taxa and address the non-monophyly of the genus Accipiter (Accipitriformes: Accipitridae)". Biological Journal of the Linnean Society: blae028. doi:10.1093/biolinnean/blae028.
Gill, Frank; Donsker, David; Rasmussen, Pamela, eds. (August 2024). "Hoatzin, New World vultures, Secretarybird, raptors". IOC World Bird List Version 14.2. International Ornithologists' Union. Retrieved 22 August 2024.