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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: ParaHoxozoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Cladus: Olfactores
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Cladus: Sarcopterygii
Cladus: Rhipidistia
Cladus: Tetrapodomorpha
Cladus: Eotetrapodiformes
Cladus: Elpistostegalia
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Cladus: Sauropsida
Classis: Reptilia
Cladus: Eureptilia
Cladus: Romeriida
Cladus: Diapsida
Cladus: Neodiapsida
Cladus: Sauria
Cladus: Archelosauria
Cladus: Archosauromorpha
Cladus: Crocopoda
Cladus: Archosauriformes
Cladus: Eucrocopoda
Cladus: Archosauria
Cladus: Avemetatarsalia
Cladus: Ornithodira
Cladus: Dinosauromorpha
Cladus: Dinosauriformes
Cladus: Dracohors
Cladus: Dinosauria
Cladus: Saurischia
Cladus: Theropoda
Cladus: Neotheropoda
Cladus: Averostra
Cladus: Tetanurae
Cladus: Avetheropoda
Cladus: Coelurosauria
Cladus: Tyrannoraptora
Cladus: Maniraptoromorpha
Cladus: Maniraptoriformes
Cladus: Maniraptora
Cladus: Pennaraptora
Cladus: Paraves
Cladus: Eumaniraptora
Cladus: Avialae
Subclassis: Aves
Cladus: Avebrevicauda
Cladus: Pygostylia
Cladus: Ornithothoraces
Cladus: Ornithuromorpha
Cladus: Carinatae
Cladus: Neornithes
Infraclassis: Neognathae
Cladus: Neoaves
Ordo: Accipitriformes

Familia: Accipitridae
Subfamilia: Gypaetinae
Genera (3 + 1†): Gypaetus - Gypohierax - Neophron – †Anchigyps

Name

Gypaetinae Vieillot, 1816
References

Vieillot, L.P. 1816. Analyse d'une nouvelle ornithologie élémentaire. 70 pp. Paris: Deterville. p. 22 "Gypaëti" BHL Reference page.

The Gypaetinae is one of two subfamilies of Old World vultures the other being the Aegypiinae. Some taxonomic authorities place the Gypaetinae within the Perninae hawks. They are presently found throughout much of Africa, Asia, and southern Europe, hence being considered "Old World" vultures, but as recently as the Late Pleistocene, they were also present in North America.[1]

A 2005 study found Eutriorchis astur to be closely related.[2]
Species
Extant genera

Subfamily Genus Common and binomial names Image Range
Gypaetinae Gypaetus Storr, 1784 Bearded vulture
Gypaetus barbatus
High mountains in southern Europe, the Caucasus, Africa, the Indian subcontinent, and Tibet
Gypaetus georgii Fossil record from the late Miocene, Spain[2]
Gypohierax Rüppell, 1836 Palm-nut vulture
Gypohierax angolensis
Forest and savannah across sub-Saharan Africa
Neophron Savigny, 1809 Egyptian vulture
Neophron percnopterus
Southwestern Europe and northern Africa to India
Neophron lolis Fossil record from the late Miocene, Spain[2]
Polyboroides A. Smith, 1829

African harrier-hawk, Polyboroides typus

Sub Saharan Africa
Madagascan harrier-hawk, Polyboroides radiatus Madagascar

Fossil genera

Genera known only from fossils include:[1]

Subfamily Genus Common and binomial names Image Range
Gypaetinae Anchigyps Anchigyps voorhiesi Late Miocene to early Pliocene of North America
Arikarornis Arikarornis macdonaldi Early Miocene of North America
Mioneophron Mioneophron longirostris Late Miocene of China
Neophrontops Neophrontops americanus Miocene to Late Pleistocene of North America
Neophrontops dakotensis
Neophrontops slaughteri
Neophrontops vallecitoensis
Neophrontops vetustus
Neogyps

(sometimes placed in the Aegypiinae)

Neogyps errans
Late Pleistocene of North America
Palaeoborus

(sometimes placed in the Aegypiinae)

Palaeoborus howardae Miocene of North America

References

Li, Zhiheng; Clarke, Julia A.; Zhou, Zhonghe; Deng, Tao (2016-10-01). "A new Old World vulture from the late Miocene of China sheds light on Neogene shifts in the past diversity and distribution of the Gypaetinae". The Auk. 133 (4): 615–625. doi:10.1642/AUK-15-240.1. ISSN 1938-4254.

Sánchez-Marco, Antonio (2022). "Two new Gypaetinae (Accipitridae, Aves) from the late Miocene of Spain" (PDF). Historical Biology. 34 (8): 1534–1543. Bibcode:2022HBio...34.1534S. doi:10.1080/08912963.2022.2053117.

Other sources
Ferguson-Lees, James; Christie, David A. (2001). Raptors of the World. Illustrated by Kim Franklin, David Mead, and Philip Burton. Houghton Mifflin. ISBN 978-0-618-12762-7. Retrieved 2011-05-26.
Grimmett, Richard; Inskipp, Carol; Inskipp, Tim (1999). Birds of India, Pakistan, Nepal, Bangladesh, Bhutan, Sri Lanka, and the Maldives. Illustrated by Clive Byers et al. Princeton University Press. ISBN 978-0-691-04910-6. OCLC 43578307.
Lerner, Heather R. L.; Mindell, David P. (November 2005). "Phylogeny of eagles, Old World vultures, and other Accipitridae based on nuclear and mitochondrial DNA" (PDF). Molecular Phylogenetics and Evolution. 37 (2): 327–346. Bibcode:2005MolPE..37..327L. doi:10.1016/j.ympev.2005.04.010. ISSN 1055-7903. PMID 15925523. Archived from the original (PDF) on 6 June 2011. Retrieved 31 May 2011.

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