Perninae

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: ParaHoxozoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Cladus: Olfactores
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Cladus: Sarcopterygii
Cladus: Rhipidistia
Cladus: Tetrapodomorpha
Cladus: Eotetrapodiformes
Cladus: Elpistostegalia
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Cladus: Sauropsida
Cladus: Reptilia
Cladus: Eureptilia
Cladus: Romeriida
Subclassis: Diapsida
Cladus: Neodiapsida
Cladus: Sauria
Cladus: Archelosauria
Cladus: Archosauromorpha
Cladus: Crocopoda
Cladus: Archosauriformes
Cladus: Eucrocopoda
Cladus: Archosauria
Cladus: Avemetatarsalia
Cladus: Ornithodira
Cladus: Dinosauromorpha
Cladus: Dinosauriformes
Cladus: Dracohors
Cladus: Dinosauria
Cladus: Saurischia
Cladus: Theropoda
Cladus: Neotheropoda
Cladus: Averostra
Cladus: Tetanurae
Cladus: Avetheropoda
Cladus: Coelurosauria
Cladus: Tyrannoraptora
Cladus: Maniraptoromorpha
Cladus: Maniraptoriformes
Cladus: Maniraptora
Cladus: Pennaraptora
Cladus: Paraves
Cladus: Eumaniraptora
Cladus: Avialae
Subclassis: Aves
Cladus: Euavialae
Cladus: Avebrevicauda
Cladus: Pygostylia
Cladus: Ornithothoraces
Cladus: Euornithes
Cladus: Ornithuromorpha
Cladus: Ornithurae
Cladus: Carinatae
Parvclassis: Neornithes
Cohors: Neognathae
Cladus: Neoaves
Ordo: Accipitriformes
Familia: Accipitridae
Subfamilia: Perninae
Genera: Aviceda - Chondrohierax – Elanoides – Hamirostra - Henicopernis - Leptodon - Lophoictinia - Pernis

Genera placement uncertain: Eutriorchis
Name

Perninae Blyth, 1851
References
Primary references

Blyth, E. 1850. Conspectus of the ornithology of India, Burma, and the Malayan Peninsular, inclusive of Sindl, Asam, Ceylon, and the Nicobar Islands. Journal of the Asiatic Society of Bengal 19: p.229–239 Scansores BHL; p.317–342 Raptores BHL; p.501–517 BHL Reference page. . p. 317

Additional references

Catanach, T.E., Halley, M.R. & Pirro, S. 2024. Enigmas no longer: using ultraconserved elements to place several unusual hawk taxa and address the non-monophyly of the genus Accipiter (Accipitriformes: Accipitridae). Biological Journal of the Linnean Society blae028. DOI: 10.1093/biolinnean/blae028 ResearchGate Reference page.

The raptor subfamily Perninae includes a number of medium-sized broad-winged species. These are birds of warmer climates, although the Pernis species (European honey buzzard and crested honey buzzard) have a more extensive range.

Several of the species in this group eat mainly insects, and the honey-buzzards are specialist feeders on wasp larvae. Reptiles are also taken by several birds in this group.

Several authorities consider Gypaetinae to be within or even synonymous with Perninae.[1][2]
Taxonomy

The cladogram of the Perninae shown below is based on a molecular phylogenetic study of the Accipitridae by Therese Catanach and collaborators that was published in 2024.[3]

Genera

Image	Genus	Living species
	Eutriorchis Sharpe, 1875	Madagascar serpent eagle Eutriorchis astur
	Chondrohierax Lesson, 1843	Hook-billed kite, Chondrohierax uncinatus Cuban kite, Chondrohierax wilsonii
	Leptodon Sundevall, 1836	Grey-headed kite, Leptodon cayanensis White-collared kite, Leptodon forbesi
	Aviceda Swainson, 1836	African cuckoo-hawk, Aviceda cuculoides Madagascar cuckoo-hawk, Aviceda madagascariensis Jerdon's baza, Aviceda jerdoni Pacific baza, Aviceda subcristata Black baza, Aviceda leuphotes
	Pernis Cuvier, 1816	European honey buzzard, Pernis apivorus Crested honey buzzard, Pernis ptilorhynchus Barred honey buzzard, Pernis celebensis Philippine honey buzzard,Pernis steerei
	Elanoides Vieillot, 1818	Swallow-tailed kite Elanoides forficatus E. f. forficatus (Linn. 1758) E. f. yetapa(Vieillot, 1818)
	Hamirostra Brown, 1846	Black-breasted buzzard Hamirostra melanosternon
	Lophoictinia Kaup, 1847	Square-tailed kite Lophoictinia isura
	Henicopernis (Lesson & Garnot, 1828)	Long-tailed honey buzzard, Henicopernis longicauda Black honey buzzard, Henicopernis infuscatus

References

Griffiths, C.S.; Barrowclough, G.F.; Groth, J.G.; Mertz, L.A. (2007). "Phylogeny, diversity, and classification of the Accipitridae based on DNA sequences of the RAG-1 exon". Journal of Avian Biology. 38 (5): 587–602. doi:10.1111/j.2007.0908-8857.03971.x.
Lerner, Heather R. L.; Mindell, David P. (2005). "Phylogeny of eagles, Old World vultures, and other Accipitridae based on nuclear and mitochondrial DNA". Molecular Phylogenetics and Evolution. 37 (2): 327–346. Bibcode:2005MolPE..37..327L. doi:10.1016/j.ympev.2005.04.010. PMID 15925523.
Catanach, T.A.; Halley, M.R.; Pirro, S. (2024). "Enigmas no longer: using ultraconserved elements to place several unusual hawk taxa and address the non-monophyly of the genus Accipiter (Accipitriformes: Accipitridae)". Biological Journal of the Linnean Society: blae028. doi:10.1093/biolinnean/blae028.