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Prunella modularis

Prunella modularis (*)

Cladus: Eukaryota
Supergroup: Opisthokonta
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Superclassis: Tetrapoda
Classis: Aves
Subclassis: Carinatae
Infraclassis: Neornithes
Parvclassis: Neognathae
Ordo: Passeriformes
Subordo: Passeri
Parvordo: Passerida
Superfamilia: Passeroidea
Familia: Prunellidae
Genus: Prunella
Species: Prunella modularis
Subspecies: P. m. euxina - P. m. fuscata - P. m. hebridium - P. m. mabbotti - P. m. meinertzhageni - P. m. modularis - P. m. obscura - P. m. occidentalis


Prunella modularis (Linnaeus, 1758)


Syst. Nat. ed.10 p.184

Vernacular names
Български: Сивогуша завирушка
Česky: Pěvuška modrá
Cymraeg: Llwyd y Gwrych
Dansk: Jernspurv
Deutsch: Heckenbraunelle
Ελληνικά : Θαμνοψάλτης (Κοινός)
English: Dunnock
Français: Accenteur mouchet
Lietuvių: Erškėtžvirblis
Nederlands: Heggenmus
Polski: Pokrzywnica
Suomi: Rautiainen
Svenska: Järnsparv
Vèneto: Moreta
Walon: Morete

The Dunnock, Prunella modularis, is a small passerine bird found throughout temperate Europe and into Asia. It is by far the most widespread member of the accentor family, which otherwise consists of mountain species. It is sometimes called the Hedge Accentor, Hedge Sparrow or Hedge Warbler.

It is a resident in the milder west and south of its range, but as an insectivore, must migrate south from the colder parts of its range. It is a bird of woodland, shrub and gardens. It builds a neat nest low in a bush or conifer, laying 3-5 unspotted blue eggs.

It is a host of the Common Cuckoo. It is likely that this association is recent, since other Cuckoo hosts have learned to discriminate between eggs, and the Cuckoo has consequently evolved eggs that match those of its host. In the case of the Dunnock, there is no resemblance, yet the Cuckoo eggs are accepted.

This is a Robin-sized bird, 13.5–14 cm in length, fairly drab in appearance, and somewhat resembling a small House Sparrow with its streaked back and general shape. It is also brownish underneath, and has a fine pointed bill. Adults have a grey head. The sexes are similarly coloured. They were introduced to New Zealand between 1867 and 1882 and now breed throughout the country including on the Chatham, Antipodes, Auckland and Campbell islands.[1]

This species makes up for its drab appearance with its breeding behaviour. Females are often polyandrous, breeding with two males at once, and thus giving rise to sperm competition. Males compete for mating access to the female, but DNA fingerprinting has shown that chicks within broods often have different fathers, depending on their success at monopolising access to the fertile female. Males try to ensure their paternity during courtship by pecking at the cloaca of the female to stimulate her to eject the sperm of other males with whom the female has recently mated.[2] Males provide parental care in proportion to their mating success, so it is not uncommon to see two males and a female provisioning nestlings at one nest. Polyandry is rare in birds, with only about 2% of species showing such a mating system; the majority are monogamous, where one male and one female breed together.

The main call is a shrill, persistent "tseep" and a high trilling note,[3] which often betray the bird's otherwise inconspicuous presence. The song is rapid, thin and tinkling, a sweet warble which can be confused with the Wren but is shorter and weaker.[3] The Wren's song incorporates repeated trill sounds but the Dunnock's does not. While the Dunnock's bubbling song is very pleasant, it has been described as being similar to a squeaky trolley wheel.


1. ^ Heather, Barrie; Rogertson, Hugh (2005). The Field Guide to the Birds of New Zealand (Revised ed.). Viking Press.
2. ^ "Polyandry, cloaca-pecking and sperm competition in dunnocks". Nature (Nature Publishing Group) (302): 334–336. March 1983. http://www.nature.com/nature/journal/v302/n5906/abs/302334a0.html.
3. ^ a b Peterson, Roger; Mountfort, Guy; Hollom, P.A.D. (1954). A Field Guide to the Birds of Britain and Europe. London: Collins.

* BirdLife International (2004). Prunella modularis. 2006. IUCN Red List of Threatened Species. IUCN 2006. www.iucnredlist.org. Retrieved on 12 May 2006. Database entry includes justification for why this species is of least concern
* Blasco-Zumeta, Javier. "Dunnock". Laboratorio Virtual Ibercaja (IberCaja Obra Social) (324): 1–4. http://www.ibercajalav.net/img/324_DunnockPmodularis.pdf.
* Davies, N.B. (1954). Dunnock Behaviour and Social Evolution. Oxford: Oxford University Press.

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