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Classification System: APG IV

Superregnum: Eukaryota
Regnum: Plantae
Cladus: Angiosperms
Cladus: Eudicots
Cladus: Core eudicots
Cladus: Asterids
Cladus: Lamiids
Ordo: Gentianales

Familia: Rubiaceae
Subfamilia: Cinchonoideae
Tribus: Chiococceae
Genera: AdolphoduckeaBadusaBikkiaCatesbaeaCeratopyxisChiococca – Coutaportla – CoutareaCoutareopsisCubanolaDuidaniaEosantheErithalisExostemaHintoniaIsidoreaMerumeaNernstiaOsaPhialanthusPortlandiaSalzmanniaSchmidtottiaScolosanthusShaferocharisSiemensiaSolenandraStrumpfiaThiolliereaThogsennia
Chiococca - Chione


Chiococceae Benth. & Hook.f., Gen. Pl. 2: 9, 21. (1873)

Type genus: Chiococca P.Browne, Civ. Nat. Hist. Jamaica 164. (1756)


Catesbaeeae Benth. & Hook.f., Gen. Pl. 2: 8, 17. (1873)
Type genus: Catesbaea L., Sp. PL. 1: 109. (1753)

Note: The transfer of Strumpfia to the monotypic tribe Strumpfieae (Cinchonoideae, Rubiaceae) is not used here (Paudyal et al., 2014).

Bentham, G. & Hooker, J.D. 1873. Genera Plantarum 2: 9, 21.
Barrabé, L., Mouly, A., Lowry II, P.P. & Munzinger, J. 2011. Reinstatement of the endemic New Caledonian genus Thiollierea Montrouz. (Rubiaceae) necessitated by the polyphyly of Bikkia Reinw. as currently circumscribed. Adansonia III, 33: 115-134. DOI: 10.5252/a2011n1a8 PDF Reference page.
Manns, U. & Bremer, B. 2010. Towards a better understanding of intertribal relationships and stable tribal delimitations within Cinchonoideae s.s. (Rubiaceae). Molecular Phylogenetics and Evolution 56(1): 21-39. DOI: 10.1016/j.ympev.2010.04.002 PDF Reference page.
Paudyal, S.K., Delprete, P.G. & Motley, T.J. 2014. Using molecular, morphological, and palynological evidence to transfer Strumpfia maritima to the monotypic tribe Strumpfieae (Cinchonoideae, Rubiaceae), and a re-delimitation of the tribe Chiococceae. Systematic Botany 39(4)-1197-1203. DOI: 10.1600/036364414X682580 Reference page.
Paudyal, S.K., Delprete, P.G., Neupane, S. & Motley, T.J. 2018. Molecular phylogenetic analysis and generic delimitations in tribe Chiococceae (Cinchonoideae, Rubiaceae). Botanical Journal of the Linnean Society 187(3): 365-396. DOI: 10.1093/botlinnean/boy029 Reference page. 2019. Chiococceae. Missouri Botanical Garden. Published on the internet. Accessed: 2019 Feb. 8.

Chiococceae is a tribe of flowering plants in the family Rubiaceae and contains about 233 species in 27 genera. Most representatives occur from southern Florida to tropical and subtropical America, except for the genera Badusa and Bikkia, which are found from the Philippines to the West Pacific, and Morierina and Thiollierea, which are native to New Caledonia.

Currently accepted names

Badusa A.Gray (3 sp)
Bikkia Reinw. ex Blume (10 sp)
Catesbaea L. (17 sp)
Ceratopyxis Hook.f. (1 sp)
Ceuthocarpus Aiello (1 sp)
Chiococca P.Browne (25 sp)
Coutaportla Urb. (3 sp)
Coutarea Aubl. (6 sp)
Cubanola Aiello (2 sp)
Eosanthe Urb. (1 sp)
Erithalis P.Browne (8 sp)
Exostema (Pers.) Rich. ex Humb. & Bonpl. (43 sp)
Hintonia Bullock (3 sp)
Isidorea A.Rich ex DC. (17 sp)
Morierina Vieill. (2 sp)
Nernstia Urb. (1 sp)
Osa Aiello (1 sp)
Phialanthus Griseb. (22 sp)
Phyllacanthus Hook.f. (1 sp)
Portlandia P.Browne (6 sp)
Salzmannia DC. (1 sp)
Schmidtottia Urb. (15 sp)
Scolosanthus Vahl (27 sp)
Shaferocharis Urb. (3 sp)
Siemensia Urb. (1 sp)
Thogsennia Aiello (1 sp)
Thiollierea Montrouz. (12 sp)


Antacanthus Rich. ex DC. = Scolosanthus
Asemnantha Hook.f. = Chiococca
Bathysograya Kuntze = Badusa
Bikkiopsis Brongn. & Gris = Bikkia
Cormigonus Raf. = Bikkia
Cutaria Brign. = Coutarea
Dolichanthera Schltr. & K.Krause = Morierina
Echinodendrum A.Rich. = Scolosanthus
Gonianthes Rich. = Cubanola
Herrera Adans. = Exostema
Lorencea Borhidi = Coutaportla
Solenandra Hook.f. = Exostema
Steudelago Kuntze = Exostema

External links

World Checklist of Rubiaceae
Manns, U., and B. Bremer. 2010. Towards a better understanding of intertribal relationships and stable tribal delimitations within Cinchonoideae s.s. (Rubiaceae). Molecular Phylogenetics and Evolution 56: 21-39
Motley, T. J., K. J. Wurdack, and P. G. Delprete. 2005. Molecular systematics of the Catesbaeeae-Chiococceae complex (Rubiaceae) - flower and fruit evolution and biogeographic implications. American Journal of Botany 92: 316-329
Paudyal, S. K., P. G. Delprete, and T. J. Motley. 2014. Using Molecular, Morphological, and Palynological Evidence to Transfer Strumpfia maritima to the Monotypic Tribe Strumpfieae (Cinchonoideae, Rubiaceae), and a Re-delimitation of the Tribe Chiococceae. Systematic Botany 39(4):1197-1203.

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