Copestylum

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Panorpida
Cladus: Antliophora
Ordo: Diptera
Subordo: Brachycera
Infraordo: Muscomorpha
Sectio: Aschiza
Superfamilia: Syrphoidea

Familia:Syrphidae
Subfamilia: Eristalinae
Tribus: Volucellini
Genus: Copestylum
Species (331): C. abdominale – C. abrupta – C. acutifrons – C. alberlena – C. albertoi – C. albifrons – C. albitarse – C. alcedo – C. alcedoides – C. alchimista – C. ambrosettii – C. amethystinum – C. anastasia – C. anna – C. apertum – C. apicale – C. apiciferum – C. apicula – C. aricia – C. astarte – C. aster – C. aureum – C. avidum – C. azureum – C. azurinum – C. barbara – C. barei – C. bassleri – C. beatricea – C. belinda – C. belizensis – C. bellulum – C. bequaerti – C. bimaculatum – C. binominatum – C. bipunctatum – C. bolivianum – C. boqueronense – C. bradleyi – C. brazilianum – C. breve – C. brevifacies – C. brevivittatum – C. bruneri – C. brunneum – C. brunnicolor – C. brunnigaster – C. bulbosum – C. caesariatum – C. calochaetum – C. camposi – C. capensis – C. carlosii – C. caudatum – C. chaetogaster – C. chaetophorum – C. chalybescens – C. chapadense – C. cinctiventre – C. circe – C. circumdatum – C. circumscriptum – C. claripenne – C. clarum – C. cockerelli – C. colombiense – C. compactum – C. comstocki – C. conabioi – C. concinnum – C. conifacium – C. contumax – C. cordiae – C. correctum – C. corumbense – C. craverii – C. crepuscularium – C. cruciatum – C. cubomaculatum – C. cupricolor – C. curiosum – C. currani – C. cyanescens – C. cyanoprora – C. delila – C. dichroicum – C. dimorphium – C. discale – C. dispar – C. dorsale – C. duida – C. elizabethae – C. emeralda – C. emilia – C. escomeli – C. eugenia – C. exeugenia – C. externum – C. flavipenne – C. flavissimum – C. flaviventre – C. florella – C. florida – C. flukei – C. fornax – C. fractum – C. fraudulentum – C. frauenfeldi – C. frontale – C. fulvicorne – C. fulvolucens – C. fulvonotatum – C. fumipenne – C. fumosum – C. furens – C. galantei – C. gelenitae – C. gertschi – C. gibberum – C. gorgon – C. granulatum – C. guianicum – C. haagii – C. hambletoni – C. hirtipes – C. hispaniolae – C. horticole – C. horvathi – C. hoya – C. hyalinipenne – C. hyalopterum – C. hydrofenestra – C. hystrix – C. imitans – C. impressum – C. inconsistens – C. infractum – C. inquisitor – C. integrum – C. intona – C. isabellina – C. joei – C. johnsoni – C. kahli – C. lacticoeruleum – C. lanei – C. latevitatum – C. latum – C. lentum – C. limbipenne – C. liriope – C. longirostre – C. louisae – C. lucilia – C. lugens – C. lumina – C. lunuliferum – C. macquarti – C. macrocephalum – C. macrorhinum – C. macula – C. maculoides – C. mamorum – C. marceli – C. marginatum – C. megacephalum – C. melleum – C. meretricias – C. metallorum – C. mexicanum – C. minimum – C. missionera – C. mocanum – C. morpho – C. musanum – C. muscarium – C. musicanum – C. mustoides – C. nautlanum – C. neosplendens – C. neotropicum – C. nigriceps – C. nigrifacies – C. nigripes – C. nigropodum – C. nigroviride – C. nitidigaster – C. notatum – C. obliquicorne – C. obscurior – C. obscuripenne – C. oestroides – C. omochroma – C. opalescens – C. opalicolor – C. opalina – C. opeostoma – C. opinator – C. ornatum – C. osburni – C. oscillans – C. otongaensis – C. pachecoi – C. pallens – C. pallidithorax – C. pallisteri – C. palmyra – C. panamense – C. panamenum – C. parana – C. parina – C. parvum – C. pectorale – C. persimile – C. pertinax – C. peruvianum – C. pica – C. pictum – C. picturatum – C. pinkusi – C. placivum – C. plaumanni – C. plorans – C. posticum – C. prasinus – C. prescutellare – C. procteri – C. profaupar – C. pseudopallens – C. pseudotachina – C. pubescens – C. pulchripes – C. pulchrum – C. punctiferum – C. punctigena – C. purpurascens – C. purpureum – C. purpuriferum – C. pusillum – C. puyarum – C. quadratum – C. rafaelanum – C. rectifacies – C. rectum – C. rhea – C. robustum – C. roraima – C. rosa – C. rospigliosii – C. rufitarse – C. rufoscutellare – C. rurale – C. saphirinum – C. sappho – C. satur – C. schwarzi – C. scintillans – C. scutellatum – C. selectum – C. sexmaculatum – C. sica – C. simile – C. smithae – C. soror – C. spectrale – C. spinigerum – C. spinithorax – C. splendens – C. squamigerum – C. sternale – C. stigmatum – C. subcoeruleum – C. submetallicum – C. subrostratum – C. sultzi – C. tacanense – C. tamaulipanum – C. tapanti – C. tapia – C. tatei – C. teffera – C. tibiale – C. tricinctum – C. trifascium – C. tripunctatum – C. trituberculatum – C. triunfense – C. tumicephalum – C. tympanitis – C. ulrica – C. umamas – C. unicolor – C. unilectum – C. vacuum – C. vagum – C. valeria – C. vampyrum – C. varians – C. varichaetum – C. variegatum – C. ventana – C. vera – C. verdigaster – C. vesicularium – C. vicinum – C. victoria – C. vierecki – C. villarica – C. viola – C. violaceum – C. virescens – C. viridana – C. viride – C. viridigaster – C. viridula – C. vitifacium – C. vitrea – C. vitripenne – C. vittatum – C. vittifacies – C. volcanorum – C. volucelloides – C. volucre – C. vulcan – C. vulta – C. willinki – C. wulpi – C. xalapensis – C. xipe – C. yowoi – C. yura

[Source: Catalogue of Life: 2013 Annual Checklist]
Name

Copestylum Macquart, 1846: Mém. Soc. R. Sci. Lille, 1844 (1845), 252.
References

Copestylum is one of the largest genera of hoverflies in the Americas.[3] It comprises more than 300 species,[4] of which only four have been found outside the Americas, having probably been introduced by the importation of cacti in which the larvae live.[3]
Contents

1 Taxonomy
1.1 Subgenera and type species
1.2 Species
2 Notes
3 References
4 External links

Taxonomy
Subgenera and type species

The following is a list of the subgenera within Copestylum, with just the type species of each subgenus listed afterwards.[4]

Subgenus Copestylum Macquart, 1846 – type Copestylum flaviventre Macquart, 1846[1]
Subgenus Phalacromyia Rondani, 1848 – type Copestylum submetallicum (Rondani, 1848)
Subgenus Glaurotricha Thomson, 1869 – type Copestylum muscarium (Thomson, 1869)
Subgenus Atemnocera Bigot, 1882a – type Copestylum scutellatum (Macquart, 1842)
Subgenus Apophysophora Williston, 1888 – type Copestylum trituberculatum Thomson, 1976
Subgenus Megametopon (= Ophromyia)[5] Giglio-Tos, 1891 – type Copestylum nasicum (Williston, 1891)
Subgenus Camerania Giglio-Tos, 1892a[Note 1] – type Copestylum macrocephalum (Giglio-Tos, 1892)
Subgenus Viereckomyia Curran, 1925b – type Copestylum gibberum (Schiner, 1868)
Subgenus Lepidopsis Curran, 1925b – type Copestylum compactum (Curran, 1925)
Subgenus Volosyrpha Shannon, 1929[Note 2] – type Copestylum rufitarse Thomson, 1976
Subgenus Volucellosia Curran, 1930d – type Copestylum fornax Townsend, 1895
Subgenus Tachinosyrphus Hull, 1936b – type Copestylum pseudotachina (Hull, 1936)

Species

C. abdominale (Wiedemann, 1930)
C. abrupta (Curran, 1925)
C. acutifrons (Curran, 1939)
C. alberlena Marcos-Garcia & Perez-Banon, 2002
C. albertoi Marcos-Garcia & Rotheray, 2007
C. albifrons (Curran, 1939)
C. albitarse (Lynch Arribalzaga, 1892)
C. alcedo (Curran, 1926)
C. alcedoides (Curran, 1939)
C. alchimista (Rondani, 1848)
C. ambrosettii (Lynch Arribalzaga, 1892)
C. amethystinum (Bigot, 1875)
C. anastasia (Hull, 1945)
C. anna (Williston, 1887) (Anna's bromeliad fly)
C. apertum (Walker, 1860)
C. apicale (Loew, 1866)
C. apiciferum (Townsend, 1895)
C. apicula (Curran, 1939)
C. aricia (Curran, 1930)
C. astarte (Hull, 1950)
C. aster (Curran, 1939)
C. aureum (Fluke, 1951)
C. avidum (Osten Sacken, 1877) (yellow-spotted bromeliad fly)
C. azureum (Philippi, 1865)
C. azurinum (Hull, 1941)
C. barbara Hancock & Marcos-Garcia, 2007
C. barei (Curran, 1925) (violet bromeliad fly)
C. bassleri (Curran, 1939)
C. beatricea (Hull, 1950)
C. belinda (Hull, 1949)
C. belizensis Sedman, 1975
C. bellulum (Williston, 1891)
C. bequaerti Curran, 1930
C. bimaculatum (Sack, 1941)
C. binominatum (Goot, 1964)
C. bipunctatum (Hull, 1941)
C. bolivianum (Hine, 1914)
C. boqueronense Rotheray & Marcos-Garcia, 2007
C. bradleyi (Curran, 1925)
C. brazilianum (Hull, 1938)
C. breve (Giglio-Tos, 1892)
C. brevifacies (Curran, 1926)
C. brevivittatum (Curran, 1930)
C. bruneri (Curran, 1939)
C. brunneum (Thunberg, 1789)
C. brunnicolor (Hull, 1938)
C. brunnigaster (Hull, 1943)
C. bulbosum (Fluke, 1951)
C. caesariatum (Williston, 1891)
C. calochaetum (Hull, 1941)
C. camposi (Curran, 1939)
C. capensis (Schiner, 1868)[5]
C. carlosii Rotheray & Hancock, 2007
C. caudatum Curran, 1927 (hairy-horned bromeliad fly)
C. chaetogaster (Hull, 1943)
C. chaetophorum (Williston, 1887)
C. chalybescens (Wiedemann, 1830)
C. chapadense (Curran, 1930)
C. cinctiventre (Curran, 1930)
C. circe (Curran, 1939)
C. circumdatum (Walker, 1857)
C. circumscriptum (Curran, 1939)
C. claripenne (Curran, 1925)
C. clarum (Hull, 1942)
C. cockerelli (Curran, 1927)
C. colombiense Thompson, 1976
C. compactum (Curran, 1925)
C. comstocki (Williston, 1887) (Comstock's bromeliad fly)
C. conabioi Marcos-Garcia & Rotheray, 2007
C. concinnum (Philippi, 1865)
C. conifacium (Hull, 1943)
C. contumax (Curran, 1939)
C. cordiae (Townsend, 1897)
C. correctum (Curran, 1927)
C. corumbense (Curran, 1930)
C. craverii (Giglio-Tos, 1892)
C. crepuscularium (Hull, 1942)
C. cruciatum (Hull, 1943)
C. cubomaculatum (Hull, 1937)
C. cupricolor (Hull, 1948)
C. curiosum (Curran, 1939)
C. currani (Fluke, 1951)
C. cyanescens (Macquart, 1842)
C. cyanoprora (Curran, 1939)
C. delila (Hull, 1950)
C. dichroicum (Giglio-Tos, 1892)
C. dimorphium (Curran, 1939)
C. discale (Curran, 1926)
C. dispar (Macquart, 1846)
C. dorsale (Wiedemann, 1830)
C. duida (Curran, 1930)
C. elizabethae Hancock & Rotheray, 2007
C. emeralda (Hull, 1944)
C. emilia (Curran, 1939)
C. escomeli (Curran, 1929)
C. eugenia (Williston, 1887)
C. exeugenia (Curran, 1953)
C. externum (Curran, 1939)
C. flavipenne (Wiedemann, 1830)
C. flavissimum (Giglio-Tos, 1892)
C. flaviventre (Macquart, 1846)
C. florella (Hull, 1944)
C. florida (Hull, 1941) (Florida bromeliad fly)
C. flukei (Curran, 1936)
C. fornax (Townsend, 1895)
C. fractum (Curran, 1926)
C. fraudulentum (Williston, 1891)
C. frauenfeldi (Schiner, 1868)[5]
C. frontale (Sack, 1941)
C. fulvicorne (Bigot, 1883)
C. fulvolucens (Walker, 1852)
C. fulvonotatum (Bigot, 1875)
C. fumipenne (Sack, 1941)
C. fumosum (Hull, 1943)
C. furens (Giglio-Tos, 1892)
C. galantei Marcos-Garcia & Rotheray, 2007
C. gelenitae Marcos-Garcia & Rotheray, 2007
C. gertschi (Curran, 1939)
C. gibberum (Schiner, 1868)[5]
C. gorgon (Hull, 1950)
C. granulatum (Hull, 1944)
C. guianicum (Hine, 1914)
C. haagii (Jaennicke, 1867) (Haag's bromeliad fly)
C. hambletoni (Fluke, 1951)
C. hirtipes (Macquart, 1850)
C. hispaniolae Thompson, 1981
C. horticole (Hull, 1943)
C. horvathi (Szilády, 1926)
C. hoya (Curran, 1947)
C. hyalinipenne (Hull, 1944)
C. hyalopterum (Giglio-Tos, 1892)
C. hydrofenestra (Hull, 1943)
C. hystrix (Giglio-Tos, 1892)
C. imitans (Curran, 1926)
C. impressum (Hull, 1949)
C. inconsistens (Curran, 1939)
C. infractum Thompson, 1981
C. inquisitor (Hull, 1943)
C. integrum (Walker, 1857)
C. intona (Curran, 1928)
C. isabellina (Williston, 1887) (Isabelle's bromeliad fly)
C. joei Rotheray & Hancock, 2007
C. johnsoni (Curran, 1925)
C. kahli (Hull, 1938)
C. lacticoeruleum (Hull, 1944)
C. lanei (Curran, 1936)
C. latevitatum (Curran, 1939)
C. latum (Wiedemann, 1830)
C. lentum Williston, 1887
C. limbipenne Williston, 1887
C. liriope (Hull, 1949)
C. longirostre (Macquart, 1846)
C. louisae Hancock & Rotheray, 2007
C. lucilia (Hull, 1950)
C. lugens (Wiedemann, 1830)
C. lumina (Hull, 1937)
C. lunuliferum (Hull, 1937)
C. macquarti (Curran, 1926)
C. macrocephalum (Giglio-tos, 1892)
C. macrorhinum (Bigot, 1875)
C. macula (Wiedemann, 1830)
C. maculoides (Curran, 1939)
C. mamorum Rotheray & Marcos-Garcia, 2007
C. marceli (Curran, 1939)
C. marginatum (Say, 1829)
C. megacephalum (Loew, 1863)
C. melleum (Jannicke, 1867)
C. meretricias (Williston, 1888)
C. metallorum (Walker, 1852)
C. mexicanum (Macquart, 1842) (Mexican cactus fly)
C. minimum (Giglio-Tos, 1892)
C. missionera (Lynch Arribalzaga, 1892)
C. mocanum (Curran, 1936)
C. morpho (Curran, 1939)
C. musanum (Curran, 1930)
C. muscarium (Thomson, 1869)
C. musicanum (Curran, 1930)
C. mustoides (Curran, 1927)
C. nautlanum (Townsend, 1897)
C. neosplendens Thompson, 1976
C. neotropicum Thompson, 1976
C. nigriceps (Schiner, 1868)[5]
C. nigrifacies (Bigot, 1875)
C. nigripes (Bigot, 1857)
C. nigropodum (Hull, 1949)
C. nigroviride (Hull, 1949)
C. nitidigaster (Hull, 1937)
C. notatum (Bigot, 1875)
C. obliquicorne (Curran, 1939)
C. obscurior (Curran, 1939)
C. obscuripenne (Lynch Arribalzaga, 1892)
C. oestroides (Hull, 1943)
C. omochroma (Giglio-Tos, 1892)
C. opalescens (Townsend, 1901)
C. opalicolor (Hull, 1943)
C. opalina (Wiedemann, 1830)
C. opeostoma (Hull, 1949)
C. opinator (Williston, 1891)
C. ornatum (Williston, 1891)
C. osburni (Hull, 1942)
C. oscillans Hancock & Rotheray, 2007
C. otongaensis Rotheray & Hancock, 2007
C. pachecoi (Curran, 1939)
C. pallens (Wiedemann, 1830)
C. pallidithorax (Hull, 1941)
C. pallisteri (Curran, 1953)
C. palmyra (Hull, 1949)
C. panamense (Curran, 1930)
C. panamenum (Curran, 1939)
C. parana (Hull, 1942)
C. parina (Fluke, 1951)
C. parvum (Rondani, 1848)
C. pectorale (Rondani, 1863)
C. persimile (Williston, 1888)
C. pertinax (Hull, 1950)
C. peruvianum (Vimmer & Soukup, 1938)
C. pica (Schiner, 1868)[5]
C. pictum (Wiedemann, 1830)
C. picturatum (Lynch Arribalzaga, 1892)
C. pinkusi (Curran, 1938)
C. placivum (Hull, 1943)
C. plaumanni (Curran, 1939)
C. plorans (Rondani, 1848)
C. posticum (Say, 1829)
C. prasinus (Schiner, 1868)[5]
C. prescutellare (Williston, 1888)
C. procteri (Curran, 1939)
C. profaupar Marinoni, 2004
C. pseudopallens Thompson, 1981
C. pseudotachina (Hull, 1936)
C. pubescens (Loew, 1861)
C. pulchripes (Bigot, 1875)
C. pulchrum (Williston, 1891)
C. punctiferum (Bigot, 1875)
C. punctigena (Hull, 1937)
C. purpurascens (Loew, 1869)
C. purpureum (Walker, 1849)
C. purpuriferum (Bigot, 1875)
C. pusillum (Macquart, 1842)
C. puyarum Rotheray & Hancock, 2007
C. quadratum (Williston, 1891)
C. rafaelanum (Townsend, 1897)
C. rectifacies Thompson, 1981
C. rectum (Wulp, 1882)
C. rhea (Hull, 1950)
C. robustum (Sack, 1941)
C. roraima (Curran, 1939)
C. rosa (Curran, 1939)
C. rospigliosii (Brethes, 1920)
C. rufitarse Thompson, 1976
C. rufoscutellare (Philippi, 1865)
C. rurale (Curran, 1939)
C. saphirinum (Bigot, 1883)
C. sappho (Hull, 1949)
C. satur (Osten Sacken, 1877) (spotted-wing bromeliad fly)
C. schwarzi Curran, 1935
C. scintillans (Hull, 1949)
C. scutellatum (Macquart, 1842)
C. selectum (Curran, 1939)
C. sexmaculatum (Palisot de Beauvois, 1819) (six-spotted bromeliad fly)
C. sica (Curran, 1953)
C. simile Giglio-Tos, 1892
C. smithae (Thompson, 1965)
C. soror (Bigot, 1883)
C. spectrale (Hull, 1942)
C. spinigerum (Wiedemann, 1830)
C. spinithorax (Lynch Arribalzaga, 1892)
C. splendens (Townsend, 1897)
C. squamigerum (Curran, 1925)
C. sternale (Curran, 1930)
C. stigmatum (Hull, 1949)
C. subcoeruleum (Rondani, 1863)
C. submetallicum (Rondani, 1848)
C. subrostratum (Rondani, 1848)
C. sultzi (Curran, 1939)
C. tacanense Rotheray & Hancock, 2007
C. tamaulipanum (Townsend, 1898)
C. tapanti Rotheray & Hancock, 2007
C. tapia Rotheray & Hancock, 2007
C. tatei (Curran, 1930)
C. teffera (Curran, 1939)
C. tibiale (Macquart, 1846)
C. tricinctum (Bigot, 1875)
C. trifascium (Walker, 1857)
C. tripunctatum (Hull, 1949)
C. trituberculatum Thompson, 1976
C. triunfense Hancock, 2007
C. trivittatum Thompson, 1988
C. tumicephalum (Hull, 1943)
C. tympanitis (Fabricius, 1805)
C. ulrica (Hull, 1950)
C. umamas Hancock & Rotheray, 2007
C. unicolor (Curran, 1925)
C. unilectum (Walker, 1860)
C. vacuum (Fabricius, 1775)
C. vagum (Wiedemann, 1830)
C. valeria (Hull, 1944)
C. vampyrum (Hull, 1942)
C. varians (Bigot, 1875)
C. varichaetum (Curran, 1925)
C. variegatum (Bigot, 1875)
C. ventana (Fluke, 1951)
C. vera (Hull, 1942)
C. verdigaster (Hull, 1943)
C. vesicularium (Curran, 1947) (irridescent bromeliad fly)
C. vicinum (Bigot, 1883)
C. victoria (Williston, 1887) (Victoria's bromeliad fly)
C. vierecki (Curran, 1925)
C. villarica (Fluke, 1951)
C. viola (Hull, 1944)
C. violaceum (Say, 1829) (purple bromeliad fly)
C. virescens (Williston, 1891)
C. viridana (Townsend, 1897)
C. viride (Williston, 1888)
C. viridigaster (Hull, 1943)
C. viridula (Walker, 1860)
C. vitifacium (Hull, 1943)
C. vitrea (Hull, 1949)
C. vitripenne (Curran, 1930)
C. vittatum Thompson, 1976 (striped bromeliad fly)
C. vittifacies (Hull, 1943)
C. volcanorum Hancock & Rotheray, 2007
C. volucelloides (Bigot, 1884)
C. volucre (Giglio-Tos, 1892)
C. vulcan (Hull, 1942)
C. vulta (Fluke, 1951)
C. willinki (Fluke, 1951)
C. wulpi (Goot, 1964)
C. xalapensis Rotheray & Marcos-Garcia, 2007
C. xipe (Hull, 1942)
C. yowoi Rotheray & Hancock, 2007
C. yura (Curran, 1930)
Notes

originally described as Temnocera megacephala (Loew)

Type-species, tibialis Shannon preoccupied so renamed rufitarse by Thompson[4]

References

Macquart, P.J.M. (1846). Diptères exotiques nouveaux ou peu connus. Supplement. [1]. Lille: Mem. Soc. R. Sci. Agric. Arts. pp. 133–364, 20 pls. Retrieved 5 December 2020.
Thompson, F. C.; Vockeroth, J. R. (May 27, 2007). Neal L. Evenhuis (ed.). "51. Family Syrphidae". Australasian/Oceanian Diptera Catalog – Web Version. Bishop Museum. Retrieved January 24, 2011.
M. A. Marcos-García; C. Pérez-Bañón (2002). "Life cycle, adult and immature stages of a new species of Copestylum (Diptera: Syrphidae) from Mexico reared from Cactaceae" (PDF). Annals of the Entomological Society of America. 95 (4): 432–440. doi:10.1603/0013-8746(2002)095[0432:LCAAIS]2.0.CO;2.
Thompson, F. C.; Vockeroth, J. R.; Sedman, Y. S. "A Catalogue of the Americas South of the United States: Family Syrphidae". São Paulo, Brasil: Museu de Zoologia. Retrieved 2 February 2011.
Giglio-Tos, Ermanno. "Ditteri del Messico". Torino, Italia: C. Clausen. Retrieved 16 February 2011.