Lethe

Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Panorpida
Cladus: Amphiesmenoptera
Ordo: Lepidoptera
Subordo: Glossata
Cladus: Coelolepida
Cladus: Myoglossata
Cladus: Neolepidoptera
Infraordo: Heteroneura
Cladus: Eulepidoptera
Cladus: Ditrysia
Cladus: Apoditrysia
Cladus: Obtectomera
Superfamilia: Papilionoidea

Familia: Nymphalidae
Subfamilia: Satyrinae
Tribus: Satyrini
Subtribus: Lethina
Genus: Lethe
Subgenera: L. (Magula)

Species: L. alberta – L. albolineata – L. andersoni – L. argentata – L. armandina – L. atkinsonia – L. baileyi – L. baladeva – L. baoshana – L. baucis – L. berdievi – L. bhairava – L. bitaensis – L. camilla – L. chandica – L. confusa – L. cybele – L. daretis – L. dataensis – L. diana – L. distans – L. drypetes – L. dura – L. dynsate – L. elwesi – L. europa – L. gelduba – L. gemina – L. gesangdawai – L. giancbozanoi – L. gianchi – L. goalpara – L. gracilis – L. gregoryi – L. gulnihal – L. hecate – L. houjiangae – L. huongii – L. insana – L. jalaurida – L. jianqingi – L. kabrua – L. kansa – L. kazuichiroi – L. kondoi – L. kinabalensis – L. konkakini – L. kouleikouzana – L. lanaris – L. langsongyuni – L. latiaris – L. leei – L. liae – L. lisuae – L. liyufeii – L. luojiani – L. maitrya – L. manzorum – L. margaritae – L. marginalis – L. mekara – L. melisana – L. minerva – L. monilifera – L. naga – L. neofasciata – L. nicetas – L. nicetella – L. nigrifascia – L. nosei – L. ocellata – L. paraprocne – L. philemon – L. philesana – L. philesanoides – L. procne – L. proxima – L. ramadeva – L. rohria – L. sadona – L. satyavati – L. scanda – L. serbonis – L. sicelis – L. siderea – L. sidonis – L. sinorix – L. sisii – L. sura – L. syrcis – L. tengchongensis – L. tingeda – L. tristigmata – L. umedai – L. unistigma – L. verma – L. vindhya – L. violaceopicta – L. visrava – L. wui – L. yoshikoae – L. yuemingae – L. yunnana – L. zhangi
Name

Lethe Hübner, [1819]

Type species: Papilio europa Fabricius, 1775
Synonyms

Tanaoptera Billberg, 1820
Zophoessa Doubleday, [1849]
Debis Doubleday, [1849]
Charma Doherty, 1886
Tansima Moore, 1881
Rangbia Moore, [1892]
Nemetis Moore, [1892]
Pegada Moore, [1892]
Kirrodesa Moore, [1892]
Placilla Moore, [1892]]
Archondesa Moore, [1892]
Choranesa Moore, [1892]
Dionana Moore, [1892]
Sinchula Moore, [1892]
Kerrata Moore, [1892]
Putlia Moore, [1892]
Harima Moore, [1892]
Magula Fruhstorfer, 1911
Hermias Fruhstorfer, 1911

References

Das, G.N., Lang, S.Y., Singh, N. & Chandra, K. 2020. Taxonomic notes on the jalaurida-subgroup of the genus Lethe (Lepidoptera, Nymphalidae, Satyrinae). Zootaxa 4789(1): 281–290. DOI: 10.11646/zootaxa.4789.1.11. Paywall. Reference page.
Huang, H. 1998. Research on the butterflies of the Namjagbarwa Region, S. E. Tibet (Lepidoptera: Rhopalocera). Neue Entomologische Nachrichten 41: 207–263. PDF. Reference page.
Huang, H., 1999: Some new butterflies from China 1. Lambillionea XCIX 4déc.: 642–676.
Huang, H. 2002. Some new satyrids of the tribe Lethini from China 1 (Lepidoptera, Satyridae). Atalanta 33(3-4): 361–372. Full article (PDF). Reference page.
Huang, H. 2014. New or little known butterflies from China (Lepidoptera: Nymphalidae et Lycaenidae). Atalanta 45(1–4): 151–162. full article (PDF). Reference page.
Huang, S.Y., Wang, M. & Fan, X.L. 2019. A new species of the genus Lethe Hübner, 1819 from Yunnan, China (Lepidoptera, Nymphalidae, Satyrinae). Zootaxa 4700(4): 464–470. DOI: 10.11646/zootaxa.4700.4.4 Paywall. Reference page.
Huang, H., Wu, C.S. & Yuan, F. 2003. Zophoessa ocellata (Poujade, 1885) and its allies in China with the description of two new species A review of the genera Lethe, Zophoessa and Neope in China - 1 (Lepidoptera, Satyrinae). Neue Entomologische Nachrichten 55: 145–158. Full article (PDF). Reference page.
Koiwaya, S. 1998. A new species of the genus Lethe from China (Lepidoptera: Satyridae). Gekkan-Mushi 325: 12–21. Reference page.
Lang, S.Y. 2014. A new species of Lethe Hübner, 1819 from W. China (Lepidoptera, Nymphalidae). Atalanta 45(1-4): 171–174. Reference page.
Lang, S.Y. 2014a. Notes on Lethe Hübner, 1819 from China, with descriptions of two new subspecies (Lepidoptera, Nymphalidae). Neue Entomologische Nachrichten 70: 299–308. Reference page.
Lang, S-Y. 2015. Lethe gregoryi Watkins (Lepidoptera, Nymphalidae, Satyrinae), a distinct species. Zootaxa 4048(4): 575–582. DOI: 10.11646/zootaxa.4048.4.8. Preview (PDF) Reference page.
Lang, S.Y. 2016c. Notes on some little known Lethe Hübner, 1819 from Nujiang and Dulongjiang, N W Yunnan, China, with descriptions of a new subspecies (Lepidoptera, Nymphalidae, Satyrinae). Atalanta 47: 216-223. Reference page.
Lang, S.Y. 2016d. Notes on some Lethe species from W. China with description of two new subspecies (Lepidoptera, Nymphalidae, Satyrinae). Atalanta 47: 341-350. Reference page.
Lang, S.Y. 2016e. Study on Lethe procne (LEECH, 1891) and Lethe paraprocne Lang & Liu, 2014 with the description of two new subspecies from SW China (Lepidoptera, Nymphalidae, Satyrinae). Atalanta 47: 230-237. Reference page.
Lang, S-Y. & Duan, S-Z. 2016. Description of a new species of Lethe (Lepidoptera, Nymphalidae, Satyrinae) from western Yunnan, China. Zootaxa 4179(2): 295–300. DOI: 10.11646/zootaxa.4179.2.10. Reference page.
Lang, S-Y. & Lamas, G. 2016. What is Lethe hyrania (Kollar, 1844) (Lepidoptera, Nymphalidae, Satyrinae)? Zootaxa 4072(3): 396–400. DOI: 10.11646/zootaxa.4072.3.10. Reference page.
Lang, S.Y. & Liu, Z.H. 2014. Descriptions of one new species and one new subspecies of the genus Lethe Hübner, 1819 from SW China (Lepidoptera, Nymphalidae). Atalanta 45(1-4): 167–170. Reference page.
Lang, S-Y. & Monastyrskii, A.L. 2016. Description of two new species of the Lethe manzorum-group (Lepidoptera, Nymphalidae, Satyrinae) from China. Zootaxa 4103(5): 453–562. DOI: 10.11646/zootaxa.4103.5.3. Reference page.
Lang, S.Y. & Wang, C.H. 2016. Description of a new species of the Lethe nigrifascia LEECH, 1890-group (Lepidoptera, Nymphalidae, Satyrinae). Atalanta 47: 224-229. Reference page.
Monastyrskii, A.L., 2004: Two new species of Nymphalidae (Satyrinae) from Vietnam (Lepidoptera: Rhopalocera). Atalanta 35 (1-2): 45–49.
Monastyrskii, A.L., 2012: New taxa and new records of butterflies from Vietnam (4) (Lepidoptera: Rhopalocera). Atalanta 43 (1/2): 156–164.
Monastyrskii, A.L. & A.L. Devyatkin, 2000: New taxa and new records of butterflies from Vietnam (Lepidoptera: Rhopalocera). Atalanta 31(3/4): 471–492.
Monastyrskii, A.L. & Devyatkin, A.L., 2003. New taxa and new records of butterflies from Vietnam 2. Atalanta 34(1-2): 75–109. PDF. Reference page.
Satoshi, K. 2011: Description of a new species of Zophoessa (Satyridae) from Southern China. Gekkan-Mushi, (481): 41–45. [in Japanese] [not seen]
Satoshi, K.; ??? 2011: Description of a new species of Zophoessa (Satyridae) from Northern Myanmar. Gekkan-Mushi, (480): 31–33. [in Japanese] [not seen]
Yamamoto, A. & Takei, S., 1993: A new subspecies of Zophoessa dataensis (Semper) (Lepidoptera, Satyridae) from Mindoro, the Philippines. Tyô to Ga, 43 (4): 237–238. Abstract and full article: [1]
Yoshino, K., 2008: New species and new subspecies of Lethe from Myanmar, China and Vietnam. Futoa 54: 9–15.
Zhai, Q. & Y.-L. Zhang, 2011: A new species and new subspecies of Lethe Hübner (Lepidoptera: Nymphalidae: Satyrinae) from China. Acta Zootaxonomia Sinica 36 (1): 45–48.

Vernacular names
日本語: ヒカゲチョウ属
References

A Check List of Butterflies in Indo-China

Lethe is a butterfly genus from the subfamily Satyrinae in the family Nymphalidae. The genus was erected by Jacob Hübner in 1819. It includes the treebrowns, woodbrowns, foresters and their relatives. The species in the genus Lethe occur in temperate-tropical southern and eastern Asia, up to Indonesia and in North America.
Lethe anthedon, northern pearly-eye
Lethe
Selected species

These 108 species belong to the genus Lethe.

Lethe albolineata Poujade, 1884 c g
Lethe andersoni Atkinson, 1871 c g
Lethe anthedon (A. Clark, 1936) i g b (northern pearly-eye)
Lethe appalachia R. Chermock, 1947 i g b (Appalachian brown)
Lethe arete Cramer, 1780 c g
Lethe argentata Leech, 1891 c g
Lethe armandina Oberthür, 1881 c g (Chinese labyrinth )
Lethe atkinsonia Hewitson, 1876 c g (small goldenfork )
Lethe baileyi South, 1913 c g
Lethe baladeva Moore, 1865 c g (treble silverstripe )
Lethe baucis Leech, 1891 c g
Lethe bhairava Moore, 1857 c g (rusty forester)
Lethe bojonia Fruhstorfer, 1913 g
Lethe brisanda de Nicéville, 1886 c g
Lethe butleri Leech, 1889 c g
Lethe butleri periscelis Fruhstorfer, 1908
Lethe callipteris Butler, 1877 c g
Lethe camilla Leech, 1891 c g
Lethe chandica Moore, 1857 c g (angled red forester)
Lethe chandica ratnacri Fruhstorfer, 1908
Lethe christophi Leech, 1891 c g
Lethe christophi hanako Fruhstorfer, 1908
Lethe confusa Aurivillius, 1897 c g (banded treebrown)
Lethe creola (Skinner, 1897) i g b (creole pearly-eye)
Lethe cybele Leech c g
Lethe cyrene Leech, 1890 c g
Lethe dakwania Tytler, 1939 c g
Lethe darena Felder, 1867 c g
Lethe daretis Hewitson, 1863 c g (Sri Lanka treebrown)
Lethe dejeani Oberthür c g
Lethe delila Staudinger, 1896 c g
Lethe diana Butler, 1866 c g
Lethe diana australis Naritomi, 1943
Lethe distans Butler, 1870 c g (scarce red forester)
Lethe dora Staudinger, 1896 c g
Lethe drypetis Hewitson, 1863 c g (Tamil treebrown)
Lethe dura Marshall, 1882 c g (scarce lilacfork)
Lethe dynsate Hewitson, 1863 c g (Ceylon forester)
Lethe dyrta Felder, 1867 c g
Lethe elwesi Moore, 1892 g
Lethe europa Fabricius, 1775 c g (bamboo treebrown)
Lethe europa pavida Fruhstorfer, 1908
Lethe eurydice (Linnaeus, 1763) i c g b (eyed brown)
Lethe fumosus Leussler, 1916 c g
Lethe gemina Leech, 1891 c g (Tytler's treebrown)
Lethe gemina zaitha Fruhstorfer, 1914
Lethe giancbozanoi Lang & Monastyrskii g
Lethe goalpara Moore, 1865 c g (large goldenfork)
Lethe gracilis Oberthür, 1886 c g
Lethe gregoryi Watkins, 1927 g
Lethe gulnihal de Nicéville, 1887 - dull forester
Lethe hecate Leech, 1891 c g (dull forester)
Lethe helena Leech, 1891 c g
Lethe helle Leech, 1891 c g
Lethe insana Kollar, 1844 c g (common forester)
Lethe insana formosana Fruhstorfer, 1908
Lethe insularis Fruhstorfer, 1911 c g
Lethe irma Evans, 1923 c g
Lethe jalaurida de Nicéville, 1880 c g (small silverfork)
Lethe kabrua Tytler, 1914 c g (Manipur goldenfork)
Lethe kanjupkula Tytler, 1914 c g
Lethe kansa Moore, 1857 c g (bamboo forester)
Lethe kinabalensis Okubo, 1979 c g
Lethe labyrinthea Leech, 1890 c g
Lethe lanaris Butler, 1877 c g
Lethe laodamia Leech, 1891 c g
Lethe latiaris Hewitson, 1863 c g (pale forester)
Lethe luteofasciata Poujade, 1884 c g
Lethe maitrya de Nicéville, 1880 c g (barred woodbrown)
Lethe manthara Felder, 1867 c g
Lethe manzorum Poujade, 1884 c g
Lethe margaritae Elwes, 1882 c g (Bhutan treebrown)
Lethe marginalis Motschulsky, 1860 c g
Lethe mataja Fruhstorfer, 1908 c g
Lethe mekara Moore, 1857 c g (common red forester)
Lethe minerva Fabricius, 1775 c g
Lethe moelleri Elwes, 1887 c g (Moeller's silverfork)
Lethe monilifera Oberthür, 1923 c g
Lethe montana g
Lethe naga Doherty, 1889 c g (Naga treebrown)
Lethe nicetas Hewitson, [1868] - yellow woodbrown
Lethe nicetella de Nicéville, 1887 - small woodbrown
Lethe nicevillei Evans, 1924 c g (small woodbrown)
Lethe nigrifascia Leech, 1890 c g
Lethe ocellata Poujade, 1885 c g (dismal mystic)
Lethe oculatissima Poujade, 1885 c g
Lethe perimede Staudinger, 1895 c g
Lethe portlandia (Fabricius, 1781) i b (southern pearly-eye)
Lethe procne Leech, 1891 c g
Lethe proxima Leech, 1892/94 c g
Lethe ramadeva De Nicéville, 1887 c g (single silverstripe)
Lethe rohria Fabricius, 1787 c g (common treebrown)
Lethe rohria daemoniaca Fruhstorfer, 1908
Lethe samio Doubleday, [1849] c g
Lethe satyavati de Nicéville, 1880 - pallid forester
Lethe satyrina Butler, 1871 c g (pallid forester)
Lethe scanda Moore, 1857 c g (blue forester)
Lethe serbonis Hewitson, 1876 c g (brown forester)
Lethe serbonis serbonis
Lethe serbonis teesta
Lethe sicelis Hewitson, 1862 c g
Lethe siderea Marshall, 1880 c g - scarce woodbrown
Lethe sidonis Hewitson, 1863 c g (common woodbrown)
Lethe sinorix Hewitson, 1863 c g (tailed red forester)
Lethe sisii Lang & Monastyrskii g
Lethe sura Doubleday, [1849] c g (lilacfork)
Lethe syrcis Doubleday, [1849] c g
Lethe titania Leech, 1891 c g
Lethe trimacula Leech, 1890 c g
Lethe tristigmata Elwes, 1887 c g (spotted mystic)
Lethe vaivarta Doherty, 1886 c g
Lethe verma Kollar, 1844 c g (straight-banded treebrown)
Lethe verma cintamani Fruhstorfer, 1848
Lethe vindhya Felder, 1859 c g (black forester)
Lethe violaceopicta Poujade, 1884 c g (Manipur woodbrown)
Lethe visrava Moore, 1865 c g (white-edged woodbrown)
Lethe yantra Fruhstorfer, 1914 c g
Lethe zachara Fruhstorfer c g

Data sources: i = ITIS,[1] c = Catalogue of Life,[2] g = GBIF,[3] b = Bugguide.net[4] [5]
References

"Lethe Report". Integrated Taxonomic Information System. Retrieved 2018-04-14.
"Browse Lethe". Catalogue of Life. Retrieved 2018-04-14.
"Lethe". GBIF. Retrieved 2018-04-14.
"Lethe Genus: Information". BugGuide. Retrieved 2018-04-14.
"Lethe Hübner, [1819]" at Markku Savela's Lepidoptera and Some Other Life Forms