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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Cladus: Pancrustacea
Superclassis: Multicrustacea
Classis: Malacostraca
Subclassis: Eumalacostraca
Superordo: Peracarida
Ordo: Isopoda
Cladus: Scutocoxifera
Subordo: Cymothoida
Superfamilia: Bopyroidea

Familia: Bopyridae
Subfamiliae: Athelginae – Hemiarthrinae – Ioninae – Orbioninae – Pseudioninae
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Overview of genera (158)

AcrobelioneAlbunioneAllathelgesAllobopyrusAllodiplophryxusAllokeponAllorbimorphusAnacepon – Anathelges – Anchiarthrus – Anisarthrus – Anisorbione – Anomophryxus – Anuropodione – Apocepon – Apophrixus – Aporobopyrina – Aporobopyroides – Aporobopyrus – Argeia – Argeiopsis – Astalione – Asymmetrione – Asymmetrorbione – Athelges – Atypocepon – Azygopleon – Balanopleon – Bathione – Bathygyge – Bopyrella – Bopyrina – Bopyrinella – Bopyrinina – Bopyrione – Bopyriscus – Bopyrissa – Bopyroides – Bopyrophryxus – Bopyrosa – Bopyrus – Cancricepon – Carcinione – Cardiocepon – Castrione – Cataphryxus – Coxalione – Cryptobopyrus – Dactylokepon – Dicropleon – Diplophryxus – Entophilus – Eophrixus – Epicepon – Epipenaeon – Epiphrixus – Eragia – Ergyne – Eriphrixus – Falsanathelges – Filophryxus – Galathocrypta – Gareia – Gigantione – Goleathopseudione – Grapsicepon – Gyge – Hemiarthrus – Hemicepon – Hemiphryxus – Heterocepon – Hypercepon – Hyperphrixus – Hypocepon – Hypoperphrixus – Hypophryxus – [[[Ione (Bopyridae)|Ione]] – Ionella – Kepon – Kolourione – Leidya – Litobopyrus – Lobocepon
Loki – Mediophrixus – Megacepon – Mesocepon – Mesophryxus – Metacepon – Metaphrixus – Minicopenaeon – Minimathelges – Miophrixus – Munidion – Nikione – Ogyridione – Onychocepon – Orbimorphus – Orbione – Orophryxus – Orthione – Ovobopyrus – Pagurion – Paguristione – Palaemonellione – Parabopyrella – Parabopyriscus – Parabopyrus – Paracepon – Paragigantione – Parapagurion – Parapenaeon – Parapenaeonella – Paraphrixus – Parapleurocrypta – Parapleurocryptella – Parargeia – Parasymmetrione – Parasymmetrorbione – Parathelges – Parione – Parionella – Parionina – Parioninella – Pauperella – Phryxus – Phyllodurus – Pleurocrypta – Pleurocryptella – Pleurocryptina – Pontobopyrus – Portunicepon – Probopyria – Probopyrinella – Probopyrione – Probopyrus – Probynia – Procepon – Progebiophilus – Propseudione – Prosynsynella – Pseudione – Pseudionella – Pseudostegias – Rhopalione – Robinione – Schizobopyrina – Scyracepon – Shiinoella – Spathione – Stegias – Stegoalpheon – Synsynella – Trapezicepon – Tylokepon – Upogebione – Upogebiophilus – Urobopyrus
Name

Bopyridae Rafinesque-Schmaltz, 1815
References

An, J., Markham, J.C. & Yu, H. 2010. Description of two new species and a new genus of bopyrid isopod parasites (Bopyridae: Pseudioninae) of hermit crabs from China. Journal of Natural History 44: 2065–2073. DOI: 10.1080/00222933.2010.488753
An, J-M., Zhao, Q-P. & Markham, J.C. 2016. Paguristione uniuropodus, a new genus and a new species of Pseudioninae infesting hermit crabs from China (Crustacea, Isopoda, Bopyridae). Zookeys 577: 43–53. DOI: 10.3897/zookeys.577.6295.Reference page.
Boyko, C.B. 2003. A new genus and species of bopyrid isopod (Crustacea, Isopoda, Bopyridae, Orbioninae) parasitic on Sicyonia (Crustacea, Decapoda, Penaeoidea) from New Caledonia. Zoosystema 25: 593–600. PDF
Boyko, C.B., Williams, J.D. 1999. A revision of Anathelges and Stegophryxus (Isopoda: Bopyridae: Athelginae), with the description of two new genera and one new species. Journal of Crustacean Biology 23: 795–813. DOI: 10.1651/C-2398
Boyko, C.B., Williams, J.D. & Shields, J.D. 2017. Parasites (Isopoda: Epicaridea and Nematoda) from ghost and mud shrimp (Decapoda: Axiidea and Gebiidea) with descriptions of a new genus and a new species of bopyrid isopod and clarification of Pseudione Kossmann, 1881. Zootaxa 4365(3): 251–301. DOI: 10.11646/zootaxa.4365.3.1. Reference page.
Boyko, C.B., Williams, J.D. & Markham, J.C. 2012. Recent and fossil Isopoda Bopyridae parasitic on squat lobsters and porcelain crabs (Crustacea: Anomura: Chirostyloidea and Galatheoidea), with notes on nomenclature and biogeography. Zootaxa 3150: 1–35. Preview Reference page.
Markham, J.C. 2009. A review of the Bopyridae (Crustacea: Isopoda) of Singapore, with the addition of four species to that fauna. Raffles Bulletin of Zoology Suppl. 22: 225–236. PDF
Page, R.D.M. 1985. Review of the New Zealand Bopyridae (Crustacea: Isopoda: Epicaridea). New Zealand journal of zoology 12: 185–212.
Yang, H-M., Sluys, R., Kawakatsu, M. & Min, G-S. 2018. New molecular sequences for two genera of marine planarians facilitate determination of their position in the phylogenetic tree, with new records for two species (Platyhelminthes, Tricladida, Maricola). ZooKeys 781: 1–17. DOI: 10.3897/zookeys.778.26324 Reference page.
Zhang, X.-Q., An, J.-M. & Li, X.-Z. 2013. Cladistic analysis of the phylogenetic relationships among genera of family Bopyridae (Crustacea, Isopoda) based on morphological characters. Acta Zootaxonomica Sinica, 38(2): 207–214.

The Bopyridae are a family of isopod crustaceans in the suborder Cymothoida. There are 1223 individual species contained in this family.[1][2] Members of the family are ectoparasites of crabs and shrimp. They live in the gill cavities or under the carapace where they cause a noticeable swelling. Fossil crustaceans have occasionally been observed to have a similar characteristic bulge.[3]
Genera

Acrobelione Bourdon, 1981
Albunione Markham & Boyko, 1999
Allathelges Kazmi & Markham, 1999
Allobopyrus Bourdon, 1983
Allodiplophryxus Markham, 1985
Allokepon Markham, 1982
Allorbimorphus Bourdon, 1976
Anacepon Nierstrasz & Brender à Brandis, 1931
Anathelges Bonnier, 1900
Anchiarthrus Markham, 1992
Anisarthrus Giard, 1907
Anisorbione Bourdon, 1981
Anomophryxus Shiino, 1937
Anuropodione Bourdon, 1967
Apocepon Nierstrasz & Brender à Brandis, 1930
Apophrixus Nierstrasz & Brender à Brandis, 1931
Aporobopyrina Shiino, 1934
Aporobopyroides Nobili, 1906
Aporobopyrus Nobili, 1906
Argeia Dana, 1853
Argeiopsis Kensley, 1974
Astalione Markham, 1975
Asymmetrione Codreanu, Codreanu & Pike, 1965
Asymmetrorbione Boyko, 2003
Athelges Gerstaeker, 1862
Atypocepon Nierstrasz & Brender à Brandis, 1931
Azygopleon Markham, 1985
Balanopleon Markham, 1974
Bathione Román-Contreras & Boyko, 2007
Bathygyge Hansen, 1897
Bopyrella Bonnier, 1900
Bopyrina Kossmann, 1881
Bopyrinella Nierstrasz & Brender à Brandis, 1925
Bopyrinina Shiino, 1933
Bopyrione Bourdon & Markham, 1980
Bopyrissa Nierstrasz & Brender à Brandis, 1931
Bopyroides Stimpson, 1864
Bopyrophryxus Codreanu, 1965
Bopyrosa Nierstrasz & Brender à Brandis, 1923
Bopyrus Latreille, 1802
Cancricepon Giard & Bonnier, 1887
Carcinione Bourdon, 1983
Cardiocepon Nobili, 1906
Castrione Lima, 1980
Cataphryxus Shiino, 1936
Coxalione Bourdon, 1977
Cryptobopyrus Schultz, 1977
Dactylokepon Stebbing, 1910
Dicropleon Markham, 1972
Diplophryxus Richardson, 1904
Discomorphus Markham, 2008
Discorsobopyrus Boyko, 2004
Entophilus Richardson, 1903
Eophrixus Caroli, 1930
Epicepon Nierstrasz & Brender à Brandis, 1931
Epipenaeon Nobili, 1906
Epiphrixus Nierstraz & Brender à Brandis, 1932
Eragia Markham, 1994
Ergyne Risso, 1816
Eriphrixus Markham, 1990
Falsanathelges Boyko & Williams, 2003
Filophryxus Bruce, 1972
Galathocrypta Román-Contreras & Soto, 2002
Gareia Bourdon & Bruce, 1983
Gigantione Kossmann, 1881
Goleathopseudione Román-Contreras, 2008
Grapsicepon Giard & Bonnier, 1887
Gyge Cornalia & Panceri, 1861
Hemiarthrus Giard & Bonnier, 1887
Hemicepon Lemos de Castro & Brasil-Lima, 1980
Hemiphryxus Bruce, 1978
Heterocepon Shiino, 1936
Hypercepon Danforth, 1972
Hyperphrixus Nierstrasz & Brender à Brandis, 1931
Hypocepon Nierstrasz & Brender à Brandis, 1930
Hypohyperphrixus Nierstrasz & Brender à Brandis, 1932
Hypophryxus Shiino, 1934
Ionella Bonnier, 1900
Kepon Duvernoy, 1840
Kolourione Markham, 1978
Leidya Cornalia & Panceri, 1861
Litobopyrus Markham, 1982
Lobocepon Nobili, 1905
Loki Markham, 1972
Mediophrixus Markham, 1990
Megacepon George, 1947
Mesocepon Shiino, 1951
Mesophryxus Bruce, 1973
Metacepon Nierstrasz & Brender à Brandis, 1931
Metaphrixus Nierstrasz & Brender à Brandis, 1931
Metathelges Nierstrasz & Brender à Brandis, 1923
Minicopenaeon Bourdon, 1981
Miophrixus Barnard, 1955
Munidion Hansen, 1897
Neophryxus Bruce, 2007
Nikione Kensley, 1974
Ogyridione Markham, 1988
Onkokepon An, Yu & Li, 2006
Onychocepon Pérez, 1921
Orbimorphus Richardson, 1910
Orbione Bonnier, 1900
Orophryxus Bruce, 1972
Orthione Markham, 1988
Ovobopyrus Markham, 1985
Ovoionella Boyko, 2004
Pagurion Shiino, 1933
Pagurocryptella Boyko & Williams, 2010
Palaemonellione Markham, 1989
Parabopyrella Markham, 1985
Parabopyriscus Markham, 1985
Parabopyrus Shiino, 1934
Paracepon Nierstrasz & Brender à Brandis, 1931
Paragigantione Barnard, 1920
Parapagurion Shiino, 1933
Parapenaeon Richardson, 1904
Parapenaeonella Shiino, 1949
Parapleurocrypta Chopra, 1923
Parapleurocryptella Bourdon, 1972
Parapseudione Nierstrasz & Brender à Brandis, 1931
Parargeia Hansen, 1897
Parasymmetrione An, Markham & Yu, 2010
Parasymmetrorbione An, Boyko & Li, 2013[4]
Parathelges Bonnier, 1900
Parione Richardson, 1910
Parionella Nierstrasz & Brender à Brandis, 1923
Parionina Nierstrasz & Brender à Brandis, 1929
Parioninella Nierstrasz & Brender à Brandis, 1930
Pauperella Nierstrasz & Brender à Brandis, 1929
Phyllodurus Stimpson, 1857
Pleurocrypta Hesse, 1865
Pleurocryptella Bonnier, 1900
Pleurocryptina Nierstrasz & Brender à Brandis, 1929
Pliophrixus Caroli, 1930
Pontobopyrus Markham, 1979
Probopyria Markham, 1985
Probopyrinella Nierstrasz & Brender à Brandis, 1929
Probopyrione Bourdon, 1983
Probopyriscus Markham, 1982
Probopyrus Giard & Bonnier, 1888
Probynia Bourdon & Bruce, 1983
Procepon Shiino, 1937
Progebiophilus Codreanu & Codreanu, 1963
Propseudione Shiino, 1933
Pseudione Kossmann, 1881
Pseudionella Shiino, 1949
Pseudostegias Shiino, 1933
Rhopalione Pérez, 1920
Robinione Boyko, Williams & Shields, 2017[5]
Schizobopyrina Markham, 1985
Scyracepon Tattersall, 1905
Shiinoella Bourdon, 1972
Stegias Richardson, 1904
Stegoalpheon Chopra, 1923
Synsynella Hay, 1917
Trapezicepon Bonnier, 1900
Tylokepon Stebbing, 1904
Upogebione Markham, 1985
Upogebiophilus Nobili, 1906
Urobopyrus Richardson, 1904

References

Chris Boyko & Marilyn Schotte (2011). M. Schotte; C. B. Boyko; N. L. Bruce; G. C. B. Poore; S. Taiti & G. D. F. Wilson (eds.). "Bopyridae". World Marine, Freshwater and Terrestrial Isopod Crustaceans database. World Register of Marine Species. Retrieved June 12, 2011.
"WoRMS taxon details, Bopyridae Rafinesque, 1815". World Register of Marine Species. 2019-06-24.
Virginia Institute of Marine Science
An, J., Bokyo, C. & Li, X. (2013). A new genus and species of parasitic isopod (Isopoda: Bopyridae: Orbioninae) from Chinese waters, with a new locality and host record for Orbione halipori Nierstrasz & Brender à Brandis, 1923. Systematic Parasitology, 84(2), 149–156.
Boyko, C. B., Williams, J. D. & Shields, J. D. (2017). Parasites (Isopoda: Epicaridea and Nematoda) from Ghost and Mud Shrimp (Decapoda: Axiidea and Gebiidea) with Descriptions of a New Genus and a New Species of Bopyrid Isopod and Clarification of Pseudione Kossmann, 1881. Zootaxa, 4365, 251–301.

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