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Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Panorpida
Cladus: Antliophora
Ordo: Diptera
Subordo: Brachycera
Infraordo: Asilomorpha
Superfamilia: Nemestrinoidea

Familia: Acroceridae
Subfamilia: Philopotinae
Genus: Helle
Species (2): H. longirostris – H. rufescens

[source: Systema Dipterorum Version 1.0. Last updated: 10 August 2010]

Helle Osten-Sacken, 1896: 16

Type species: Acrocera longirostris Hudson, 1892, by monotypy.
Primary references

Osten-Sacken, C.R. 1896. A new genus of Cyrtidae (Dipt.) from New Zealand. Entomologist's Monthly Magazine (2) 7: 16–18. BHL.

Additional references

Brunetti, E.A. 1926. New and little-known Cyrtidae (Diptera). Annals and Magazine of Natural History Series 9 18(107): 561–606. DOI: 10.1080/00222932608633552 Paywall. BUGZ. Reference page.
Emberson, R.; Syrett, P.; Groenteman, R. 2011: Bankside Scientific Reserve - an important refuge for Canterbury's dryland insect fauna. Weta, 41: 4–26.
Gillung, J.P. & Winterton, S.L. 2011. New genera of philopotine spider flies (Diptera, Acroceridae) with a key to living and fossil genera. ZooKeys 127: 15–27. DOI: 10.3897/zookeys.127.1824 Open access. Reference page.
Paramonov, S.J. 1955. New Zealand Cyrtidae (Diptera) and the problem of the Pacific Island fauna. Pacific science 9(1): 16–25. hdl: 10125/8835 Open access
Schlinger, E.I. 1961. A review of the Acroceridae of Madagascar (Diptera). Mémoires de l'Institut Scientifique de Madagascar (E) 12: 257–267. PDF
Winterton, S.L. 2012. Review of Australasian spider flies (Diptera, Acroceridae) with a revision of Panops Lamarck. ZooKeys 172: 7–75. DOI: 10.3897/zookeys.172.1889 Open access. Reference page.
Winterton, S.L., Wiegmann, B.M. & Schlinger, E.I. 2007. Phylogeny and Bayesian divergence time estimations of small-headed flies (Diptera: Acroceridae) using multiple molecular markers. Molecular phylogenetics and evolution 43(3): 808–832. DOI: 10.1016/j.ympev.2006.08.015

Helle is a genus of small-headed flies. It is endemic to New Zealand.[2] The genus is named after Helle, the daughter of Athamas, from Greek mythology.[1]

Helle longirostris (Hudson, 1892)
Helle rufescens Brunetti, 1926[3]


Osten Sacken, C.R. (1896). "A new genus of Cyrtidae (Dipt.) from New Zealand". Entomologist's Monthly Magazine. 32: 16–18.
Winterton, S. L. (2012). "Review of Australasian spider flies (Diptera, Acroceridae) with a revision of Panops Lamarck". ZooKeys (172): 7–75. doi:10.3897/zookeys.172.1889. PMC 3307363. PMID 22448114.
Brunetti, E. (1926). "New and little-known Cyrtidæ (Diptera)". Annals and Magazine of Natural History. 9. 18 (107): 561–606. doi:10.1080/00222932608633552.

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