Hydrochus

Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Ordo: Coleoptera
Subordo: Polyphaga
Infraordo: Staphyliniformia
Superfamilia: Hydrophiloidea

Familia: Hydrochidae
Genus: Hydrochus (Leach)
Species: H. aequalis – H. aljibensis – H. angusi – H. angustatus – H. battjai – H. beeneni – H. bicolor – H. brevis – H. brevitarsis – H. brianbrowni – H. bruggei – H. callosus – H. choennii – H. chubu – H. coeneni – H. crenatus – H. currani – H. desenderi – H. dewnaraini – H. drechseli – H. elongatus – H. excavatus – H. falsus – H. flavipennis – H. foveatus – H. grandicollis – H. granulatus – H. ibericus – H. ignicollis – H. inaequalis – H. interruptus – H. jaechi – H. japonicus – H. jenniferiduae – H. jiawanae – H. johannapietersenae – H. kirgisicus – H. lacustris – H. laferi – H. matthewsi – H. megaphallus – H. neosquamifer – H. nitidicollis – H. nodulifer – H. nooreinus – H. obscurus – H. octocarinatus – H. opacus – H. pajnii – H. piroei – H. pseudosquamifer – H. ramcharani – H. rattanae – H. rhytipterus – H. rishi – H. roberti – H. rufipes – H. rugosus – H. satishanandi – H. scabratus – H. schereri – H. schillhammeri – H. schoenmanni – H. simplex – H. smaragdineus – H. soekhnandanae – H. squamifer – H. subcupreus – H. tariqi – H. vagus – H. vanbergehenegouweni – H. verae – H. wewalkai – H. yadavi
...
Unaccepted species (1): H. spangleri
Name

Hydrochus Leach, 1817
Synonyms

Kiransus Makhan, 1994
Amrishius Makhan, 1998
Deepakius Makhan, 1998
Rishwanius Makhan, 1998
Rishihydroius Makhan, 2001
Authority for synonymy: Short & Hebauer, 2006: 317 ('syn.n.')
Soesilius Makhan, 2001
Authority for synonymy: Short & Hebauer, 2006: 317 ('syn.n.')
Aschius Makhan, 2004
Authority for synonymy: Short & Hebauer, 2006: 317 ('syn.n.')
Satishius Makhan, 2004
Authority for synonymy: Short & Hebauer, 2006: 317 ('syn.n.')

Type species

Amrishius Makhan: Hydrochus lacustris Nietner, 1857, by original designation
Deepakius Makhan: Deepakius amrishi Makhan, 1998, by original designation
Rishwanius Makhan: Hydrochus soesilae Makhan, 1994, by original designation

References

Castro, A.; Delgado, J.A. 1999: Hydrochus aljibensis sp.n., una nueva especies del sur de la Peninsula Iberica (Coleoptera, Hydrochidae). Boletín de la Asociación española de Entomología, 23(1–2): 25–28. ISSN: 0210-8984 PDF
Hidalgo-Galiana, A.; Jäch, M.A.; Ribera, I. 2010: Hydrochus farsicus sp.n. from Iran and notes on other Palearctic species of the genus (Coleoptera: Hydrophiloidea: Hydrochidae). Zootaxa, 2344: 61–64. Preview PDF
Hidalgo-Galiana, A.; Ribera, I. (in press): Late Miocene diversification of the genus Hydrochus (Coleoptera, Hydrochidae) in the west Mediterranean area. Molecular phylogenetics and evolution, DOI: 10.1016/j.ympev.2011.01.018
Makhan, D. 1992a. Twelve new Hydrochus-species from South America (Coleoptera: Hydrophilidae). Phegea 20(2): 95–103. full article (PDF). Reference page.
Makhan, D. 1993: Nine new Hydrochus species from South and Central America (Coleoptera: Hydrophilidae). Annales Historico-Naturales Musei Nationalis Hungarici, 85: 65–70.
Makhan, D. 1994: Thirty-five new Hydrochus species from the Old and the New World (Coleoptera: Hydrophilidae). Annales Historico-Naturales Musei Nationalis Hungarici, 86: 29–42.
Makhan, D. 1994: Descriptions of a new species in a new genus, Kiransus, and five new species of Hydrochus (Coleoptera, Hydrophilidae). Zoological studies, 33(2): 160–164.
Makhan, D. 1995. Descriptions of ten new species of Hydrochus from different parts of the World (Coleoptera: Hydrochidae). Phegea 23(4): 187–193. full article (PDF). Reference page.
Makhan, D. 1995: Descriptions of six new species of Hydrochus from South and North America (Coleoptera, Hydrochidae). Zoological studies, 34(1): 18–20.
Makhan, D. 1996: Descriptions of three new species of Hydrochus (Coleoptera: Hydrochidae). Phegea, 24(4): 183–185.
Makhan, D. 1998: Three new genera and two new species of Hydrochidae from different parts of the World (Coleoptera). Annales Historico-Naturales Musei Nationalis Hungarici, 90: 139–150.
Makhan, D. 2001: A new genus Rishihydroius and three new Hydrochus species (Coleoptera: Hydrochidae) from Africa. Bangladesh journal of zoology, 29(1): 83–95.
Makhan, D. 2001: A new genus, Soesilius, and a new species of Hydrochidae (Coleoptera) from America. Russian entomological journal, 10(4): 389–393.
Makhan, D. 2004: Hydrochidae of the world, Dryopidae and Hydrophilidae (Coleoptera). Calodema, 2: 11–26.
Makhan, D. 2004: Two new Hydrochidae (Coleoptera) species from the world. Acta Biologica Universitatis Daugavpiliensis, 4(2): 107–109.
Makhan, D. 2005: Hydrochus brianbrowni sp. nov. (Coleoptera: Hydrochidae), a new American species. Baltic journal of coleopterology, 5(1): 63–65.
Makhan, D.; Hawkeswood, T.J.; Khani, M.; Ezzatpanah, S. 2012: On the synonymy of Hydrochus spangleri Hellman, 2003 (Coleoptera: Hydrochidae) with a checklist of the species of Rishihydroius Makhan, 2001. Calodema, 196: 1–6.
Makhan, D.; Khani, M.; Ezzatpanah, S. 2012: The genera Hydrochus Leach and Rishihydroius Makhan in Turkey (Coleoptera: Hydrochidae). Calodema, 197: 1–6.
Makhan, D.; Khani, M.; Ezzatpanah, S. 2012: The genus Deepakius Makhan, 1998 from Africa and Madagascar (Coleoptera: Hydrochidae) with a key to species. Calodema, 198: 1–6.
Oliva, A. 2000: Hydrochus synonymies. Latissimus, 12: 14. ISSN: 0966-2235
Short, A.E.Z.; Hebauer, F. 2006: World Catalogue of Hydrophiloidea – additions and corrections, 1 (1999–2005) (Coleoptera). Koleopterologische rundschau, 76: 315–359. PDF
Steiner, W.E., Staines, C.L., McCann, J.M. & Hellman, J.L. 2003. The Seth Forest Water Scavenger Beetle, a New Species of Hydrochus (Coleoptera: Hydrophiloidea: Hydrochidae) from the Chesapeake-Delmarva Region. The Coleopterists Bulletin 57(4): 433–443. DOI: 10.1649/581 Paywall, JSTOR Hybrid open access journal. Reference page.
Valladares, L.F.; Díaz, J.A.; Delgado, J.A. 1999: Hydrochus ibericus sp.n. (Coleoptera: Hydrochidae) from the Iberian Peninsula. Aquatic insects, 21(2): 81–87. DOI: 10.1076/aqin.21.2.81.4529
Watts, C.H.S. 1999: Revision of Australian Hydrochus (Coleoptera: Hydrochidae). Records of the South Australian Museum, 32(1): 1–43. PDF
Worthington, R.J.; Hellman, J.L.; Lago, P.K. 2016: Hydrochidae (Coleoptera) of Mississippi. Transactions of the American Entomological Society, 142, 167–213. PDF

Links

Atlas of family Hydrochidae of Russia
ION
Nomenclator Zoologicus

Hydrochus is a genus of water scavenger beetles in the monogeneric family Hydrochidae, sometimes placed in the Hydrophilidae as a subfamily Hydrochinae. The family Hydrochidae includes about 180 species, all of which are placed in the genus Hydrochus. Hydrochus has also been used for a fly genus in the family Dolichopodidae. This junior homonym is a junior subjective synonym of the genus Rhaphium.

Beetles in the genus Hydrochus are found in slow moving streams or stagnant water bodies. The club of the 7 segmented antenna consists of three segments with a cup-like basal segment. The number of tarsi on the legs are usually 5-5-5 or 4-4-4 (a tiny basal segment can be hard to see). The pronotum narrows towards the rear but is narrower than the base of the elytra. They walk around inside water or swim weakly in search of invertebrate prey. They hold air under the elytra. The larvae are also aquatic and predatory. An African species was formerly described under the genus Kiransus, but along with several other African species, they are now placed in Hydrochus.[1] Hydrochids are considered to be an older branch of the Hydrophiloidea. Hydrochids have been suggested as a sister group of the Spercheidae+Hydrophilidae but the relationships are unclear. Molecular phylogenetic comparisons suggest that they are related as a sister group to Helophorus+Georissus.[2]

An extinct member of the family, Ponohydrochus is known from the Early Cretaceous (Hauterivian) Khasurty locality in Russia.[3]
Hydrochus P1310412a.jpg
Hydrochus P1120860a.jpg
Species

More than 180 species have been described in the genus. The following list includes some of the species in the genus:[4]

Hydrochus aljibensis Castro & Delgado, 1999 g
Hydrochus angusi Valladares, 1988 g
Hydrochus angustatus Germar, 1824 g
Hydrochus argutissimus Perkins, 2019
Hydrochus aschnaae (Makhan, 1994)
Hydrochus basilaris Perkins, 2019
Hydrochus bicarinatus Perkins, 2019
Hydrochus bituberculatus Perkins, 2019
Hydrochus brevis (Herbst, 1793) g
Hydrochus brevitarsis Knisch, 1922 i c g
Hydrochus brianbrowni Makhan, 2005 i c g
Hydrochus callosus LeConte, 1855 i c g
Hydrochus collaris Perkins, 2019
Hydrochus conjunctus Perkins, 2019
Hydrochus corruscans Perkins, 2019
Hydrochus crenatus (Fabricius, 1792) g
Hydrochus currani Brown, 1929 i c g b
Hydrochus debilis Sharp, 1882 i c g
Hydrochus denarius Perkins, 2019
Hydrochus drakei Knisch, 1921 g
Hydrochus elongatus (Schaller, 1783) g
Hydrochus ensifer Hansen, 1998
Hydrochus excavatus LeConte, 1855 i c g b
Hydrochus falsus Hellman in Worthington, Hellman, and Lago 2016[5]
Hydrochus farsicus g
Hydrochus flavipennis Küster, 1852 g
Hydrochus formosus Perkins, 2019
Hydrochus foveatus Haldeman, 1852 i c g
Hydrochus grandicollis Kiesenwetter, 1870 g
Hydrochus granulatus Blatchley, 1910 i c g b
Hydrochus hellmani Perkins, 2019
Hydrochus ibericus Valladares, Diaz & Delgado, 1999 g
Hydrochus ignicollis Motschulsky, 1860 g
Hydrochus inaequalis LeConte, 1855[5]
Hydrochus jaechi Makhan, 1995 i c g
Hydrochus japonicus Sharp, 1873 g
Hydrochus jiawanae Makhan, 1996 i c g
Hydrochus kellymilleri Perkins, 2019
Hydrochus kirgisicus Motschulsky, 1860 g
Hydrochus lachmoni Makhan, 1996
Hydrochus leei Perkins, 2019
Hydrochus lobatus Perkins, 2019
Hydrochus mauriciogarciai Perkins, 2019
Hydrochus megaphallus van Berge Henegouwen, 1988 g
Hydrochus minimus Blatchley, 1919 i c g
Hydrochus neosquamifer Smetana, 1988 i c g
Hydrochus nigeriensis Hansen, 1998
Hydrochus niloticus Sharp, 1903
Hydrochus nitidicollis Mulsant, 1844 g
Hydrochus nodulifer Reitter, 1897
Hydrochus nooreinus Berge Henegouwen & Sainz-Cantero, 1992 g
Hydrochus obscurus Sharp, 1882 i c g
Hydrochus octocarinatus Hochhuth, 1871 g
Hydrochus pajnii Makhan, 2000 i c g
Hydrochus pallipes Chevrolat, 1863 i c g
Hydrochus pictus Perkins, 2019
Hydrochus pseudosecretus Oliva, 1996
Hydrochus pseudosquamifer Miller, 1965 i c g b
Hydrochus pupillus Orchymont, 1939 g
Hydrochus ramcharani Makhan, 1992
Hydrochus roomylae Makhan, 2001 i c g
Hydrochus rishwani Makhan, 1994
Hydrochus rufipes Melsheimer, 1844 i c g b
Hydrochus rugosus Mulsant, 1844 i c g b
Hydrochus sagittarius Perkins, 2019
Hydrochus scabratus Mulsant, 1844 i c g b
Hydrochus schereri Makhan, 1995 i c g
Hydrochus setosus Leech, 1948 i c g
Hydrochus shorti Perkins, 2019
Hydrochus simplex Leconte, 1851 i c g b
Hydrochus soekhnandanae Makhan 1992
Hydrochus smaragdineus Fairmaire, 1879 g
Hydrochus spangleri Hellman in Steiner, Staines, McCann and Hellman, 2003 i c g
Hydrochus squamifer LeConte, 1855 i c g b
Hydrochus subcupreus Randall, 1838 i c g b
Hydrochus tariqi Ribera, Hernando & Aguilera, 1999 g
Hydrochus tarsalis Chevrolat, 1863 i c g
Hydrochus vagus LeConte, 1852 i c g
Hydrochus tuberculatus Hansen, 1998
Hydrochus variabiloides Perkins, 2019
Hydrochus variolatus LeConte, 1851 i c g
Hydrochus yadavi Makhan, 2000 i c g

Data sources: i = ITIS,[6] c = Catalogue of Life,[7] g = GBIF,[8] b = Bugguide.net[9]
References

Hansen, M. (1998). "The systematic status of Kiransus Makhan, 1994, with the description of three new species of Hydrochus Leach, 1817 (Coleoptera: Hydrochidae)". Insect Systematics & Evolution. 29 (2): 223–232. doi:10.1163/187631298x00302.
Short, Andrew Edward Z.; Fikáček, Martin (2013). "Molecular phylogeny, evolution and classification of the Hydrophilidae (Coleoptera): Classification of the Hydrophilidae". Systematic Entomology. 38 (4): 723–752. doi:10.1111/syen.12024.
"The first remarkable fossil Hydrochidae (Coleoptera) from the Lower Cretaceous (Hauterivian) of the Buryatia Republic, Russia". Cretaceous Research. 123: 104795. 2021-07-01. doi:10.1016/j.cretres.2021.104795. ISSN 0195-6671.
Hidalgo-Galiana, A.; Ribera, I. (2011). "Late Miocene diversification of the genus Hydrochus (Coleoptera, Hydrochidae) in the west Mediterranean area". Molecular Phylogenetics and Evolution. 59 (2): 377–385. doi:10.1016/j.ympev.2011.01.018. PMID 21354316.
Worthington, Reese J.; Hellman, John L.; Lago, Paul K. (2016). "Hydrochidae (Coleoptera) of Mississippi". Transactions of the American Entomological Society. 142 (2): 167–213. doi:10.3157/061.142.0206. S2CID 88919765.
"Hydrochus Report". Integrated Taxonomic Information System. Retrieved 2018-04-22.
"Browse Hydrochus". Catalogue of Life. Retrieved 2018-04-22.
"Hydrochus". GBIF. Retrieved 2018-04-22.
"Hydrochus Genus: Information". BugGuide.net. Retrieved 2018-04-22.