Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: ParaHoxozoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Cladus: Olfactores
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Cladus: Sarcopterygii
Cladus: Rhipidistia
Cladus: Tetrapodomorpha
Cladus: Eotetrapodiformes
Cladus: Elpistostegalia
Superclassis: Tetrapoda
Cladus: Batrachomorpha
Classis: Amphibia
Subclassis: Lissamphibia
Superordo: Batrachia
Cladus: Salientia
Ordo: Anura
Familia: Ceratobatrachidae
Subfamiliae (3): Alcalinae – Ceratobatrachinae – Liuraninae
Name
Ceratobatrachidae Boulenger, 1884: 212
Type genus: Ceratobatrachus Boulenger, 1884, by original designation and monotypy.
Synonymy
Ceratobatrachinae — Gadow, 1901
Cornuferinae Noble, 1931
Platymantinae Savage, 1973
Ceratobatrachini — Dubois, 1981
Liuraninae Fei, Ye & Jiang, 2010
Alcalinae Brown, Siler, Richards, Diesmos & Cannatella, 2015: 142
References
Primary references
Boulenger, G.A. 1884. Diagnoses of new reptiles and batrachians from the Solomon Islands, collected and presented to the British Museum by H. B. Guppy, Esq., M.B., H.M.S. “Lark”. Proceedings of the Zoological Society of London 1884: 210–213. BHL
Additional references
Fei, L., Ye, C.-Y. & Jiang, J.-P. 2010. Phylogenetic systematics of Ranidae. Acta Herpetologica Sinica 12: 1–43. Reference page.
Brown, R.M., Siler, D.S., Richards, S.J., Diesmos, A.C. & Cannatella, D.C. 2015. Multilocus phylogeny and a new classification for Southeast Asian and Melanesian forest frogs (family Ceratobatrachidae). Zoological journal of the Linnean Society 174(1): 130–168. DOI: 10.1111/zoj.12232 Paywall Reference page.
Yan, F., Jiang, K., Wang, K., Jin, J.-Q., Suwannapoom, C., Li, C., Vindum, J.V., Brown, R.M. & Che, J. 2016. The Australasian frog family Ceratobatrachidae in China, Myanmar and Thailand: discovery of a new Himalayan forest frog clade. Zoological Research 37(1): 7–14. DOI: 10.13918/j.issn.2095-8137.2016.1.7 Reference page.
Vernacular names
日本語: ハナトガリガエル科
The Ceratobatrachidae are a family of frogs[1][2] found in the Malay Peninsula, Borneo, the Philippines, Palau, Fiji, New Guinea, and the Admiralty, Bismarck, and Solomon Islands.[1]
Taxonomy
Ceratobatrachidae was formerly treated as a subfamily (i.e., Ceratobatrachinae) in the family Ranidae (true frogs), but have now been re-classified as a separate family. The following genera are recognised:[1]
Subfamily Alcalinae Brown, Siler, Richards, Diesmos, and Cannatella, 2015
Alcalus (5 species)
Subfamily Ceratobatrachinae Boulenger, 1884
Cornufer Tschudi, 1838 (> 50 species)
Platymantis Günther, 1858 (> 30 species)
Subfamily Liuraninae Fei, Ye, and Jiang, 2010
Liurana Dubois, 1987 (4 species)
Formerly, the following genera were also recognized in the family Ceratobatrachidae, but have now been merged into the genera above.
Batrachylodes Boulenger, 1887 (8 species)
Palmatorappia Ahl, 1927 (1 species)
Ceratobatrachus Boulenger, 1884 (1 species)
Discodeles Boulenger, 1918 (5 species)
Distribution
Ceratobatrachidae is distributed across Island Southeast Asia,[3] as well as in the Eastern Himalayas.
Genus Liurana
Eastern Himalayas: 4 species
Genus Alcalus
Borneo: 2 species (Alcalus baluensis and Alcalus rajae)
Palawan: 1 species (Alcalus mariae)
Genus Platymantis
Philippines:
Genus Cornufer
Palau: 1 species
Maluku: 3–4 species
New Guinea: 6–8 species
Bismarck Archipelago: 18–22 species
Solomon Islands: 20–25 species
Fiji: 2 species
Life history
All Ceratobatrachidae lay eggs outside of water and undergo direct development where eggs hatch directly into froglets, without free-living tadpole stages.[4]
References
Frost, Darrel R. (2014). "Ceratobatrachidae Boulenger, 1884". Amphibian Species of the World: an Online Reference. Version 6.0. American Museum of Natural History. Retrieved 22 February 2014.
"Ceratobatrachidae Boulenger, 1884". Integrated Taxonomic Information System.
Brown, Rafe M.; Siler, Cameron D.; Richards, Stephen J.; Diesmos, Arvin C.; Cannatella, David C. (2015). "Multilocus phylogeny and a new classification for Southeast Asian and Melanesian forest frogs (family Ceratobatrachidae)". Zoological Journal of the Linnean Society. 174 (1): 130–168. doi:10.1111/zoj.12232.
Fuiten, Allison Marie (2012). Skeletal Variation in Melanesian Forest Frogs (Anura: Ceratobatrachidae). M.A. thesis , University of Kansas. p. 69. Retrieved 22 February 2014.
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