Clangula hyemalis

Clangula hyemalis, Photo: Michael Lahanas

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Cladus: Sarcopterygii
Cladus: Rhipidistia
Cladus: Tetrapodomorpha
Cladus: Eotetrapodiformes
Cladus: Elpistostegalia
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Classis: Reptilia
Cladus: Eureptilia
Cladus: Romeriida
Subclassis: Diapsida
Cladus: Sauria
Infraclassis: Archosauromorpha
Cladus: Crurotarsi
Divisio: Archosauria
Cladus: Avemetatarsalia
Cladus: Ornithodira
Subtaxon: Dinosauromorpha
Cladus: Dinosauriformes
Cladus: Dracohors
Cladus: Dinosauria
Cladus: Saurischia
Cladus: Eusaurischia
Subordo: Theropoda
Cladus: Neotheropoda
Cladus: Averostra
Cladus: Tetanurae
Cladus: Avetheropoda
Cladus: Coelurosauria
Cladus: Tyrannoraptora
Cladus: Maniraptoromorpha
Cladus: Maniraptoriformes
Cladus: Maniraptora
Cladus: Pennaraptora
Cladus: Paraves
Cladus: Eumaniraptora
Cladus: Avialae
Infraclassis: Aves
Cladus: Avebrevicauda
Cladus: Pygostylia
Cladus: Ornithothoraces
Cladus: Ornithuromorpha
Cladus: Carinatae
Parvclassis: Neornithes
Cohors: Neognathae
Cladus: Pangalloanserae
Cladus: Galloanseres
Ordo: Anseriformes

Familia: Anatidae
Subfamilia: Merginae
Genus: Clangula
Species: Clangula hyemalis
Name

Clangula hyemalis (Linnaeus, 1758)
Synonyms

Anas hyemalis (protonym)

References

Linnaeus, C. 1758. Systema Naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Editio Decima, Reformata. Tomus I. Holmiæ (Stockholm): impensis direct. Laurentii Salvii. 824 pp. DOI: 10.5962/bhl.title.542 BHL p. 126 BHL Reference page.

Vernacular names
aragonés: Guite blanco
العربية: بط طويل الذنب
asturianu: Coríu Falampiu
azərbaycanca: Moryanka
башҡортса: Эяла
беларуская: Маранка
български: Ледена потапница
brezhoneg: Morhouad-kerregi
català: Ànec glacial
kaszëbsczi: Kôłpiónka
čeština: Hoholka lední
Cymraeg: Hwyaden Gynffon-hir
dansk: Havlit
Deutsch: Eisente
Ελληνικά: χιονόπαπια
English: Long-tailed Duck
Esperanto: Longvosta klangulo
español: Pato havelda
eesti: Aul
euskara: Izotz-ahate
فارسی: اردک دم‌دراز
suomi: Alli
føroyskt: Ógvella
Nordfriisk: Isfögel
français: Harelde Boréale
Frysk: Iisein
Gaeilge: Lacha earrfhada
Gàidhlig: Eun buchainn
galego: Ànec glacial
Gaelg: Laagh Lheeah
עברית: ברווז קרח
hrvatski: Patka ledara
magyar: Jegesréce
հայերեն: Բադ ծովային
Iñupiatun: Aaqhaaliq
íslenska: Hávella
italiano: Moretta codona
ᐃᓄᒃᑎᑐᑦ / inuktitut: Aa'aangiq
日本語: コオリガモ
ქართული: მეზღვია იხვი
қазақша: Ұшқыр үйрек
kalaallisut: Alleq
한국어: 바다꿩
кыргызча: Түндүк өрдөк
lietuvių: Ledinė antis
latviešu: Kākaulis
македонски: Долгоопашеста патка
монгол: Мөнгөлөг шунгаахай
नेपाली: लामपुच्छ्रे हाँस
Nederlands: IJseend
norsk nynorsk: Havelle
norsk: Havelle
polski: Lodówka
پنجابی: لمی پوچھل آلی بطخ
português: Pato-de-cauda-afilada
rumantsch: Anda da glatsch
русский: Морянка
саха тыла: Ээбиллэ
davvisámegiella: Haŋŋá
slovenčina: Ľadovka dlhochvostá
slovenščina: Zimska raca
shqip: Rosa e ngricave
српски / srpski: Ledenjarka - Ледењарка
svenska: Alfågel
ไทย: เป็ดหางยาว
Türkçe: Telkuyruk
українська: Морянка
vepsän kel’: All'
Tiếng Việt: Vịt đuôi dài
中文: 长尾鸭

The long-tailed duck (Clangula hyemalis) or coween,[2] formerly known as the oldsquaw, is a medium-sized sea duck that breeds in the tundra and taiga regions of the arctic and winters along the northern coastlines of the Atlantic and Pacific Oceans. It is the only member of the genus Clangula.
Taxonomy

The long-tailed duck was formally described by the Swedish naturalist Carl Linnaeus in 1758 in the tenth edition of his Systema Naturae. He placed it with all the other ducks in the genus Anas and coined the binomial name Anas hyemalis.[3] Linnaeus cited the English naturalist George Edwards's description and illustration of the "Long-tailed duck from Hudson's-Bay" that had been published in 1750 in the third volume of his A Natural History of Uncommon Birds.[4]

This duck is now the only species placed in the genus Clangula; the genus was introduced in 1819 by the English zoologist William Leach to accommodate the long-tailed duck, in an appendix on species to John Ross's account of his voyage to look for the Northwest Passage.[5][6][7] The genus name Clangula is a diminutive of the Latin clangere, meaning "to resound". The specific epithet hyemalis, also Latin, means "of winter".[8] The species is considered to be monotypic – no subspecies are recognised.[7]

In North American English it is sometimes called oldsquaw, though this name has fallen out of favour. In 2000, the American Ornithologists' Union (AOU) formally adopted the name long-tailed duck, in response to petitioning by a group of biologists who feared that the former name would be offensive to Native American tribes whose help was required for conservation efforts.[a] The AOU stated that "political correctness" alone was not sufficient to justify changing a long-standing name, but in this case decided to make the change because doing so would "conform with English usage in other parts of the world".[10]

An undescribed congener is known from the Middle Miocene Sajóvölgyi Formation (Late Badenian, 13–12 Mya) of Mátraszőlős, Hungary.[11]
Distribution

Long-tailed ducks breed on tundra across northern Eurasia (in Russian Siberia, Kamchatka, and Karelia, for example), the Faroe Islands, Finland, parts of southern Greenland, Iceland, Norway, as well as across northern North America (Alaska and northern Canada).

In winter, they are found on and near large bodies of seawater, such as the Northern Pacific Ocean, the North Atlantic Ocean, Hudson Bay and the American Great Lakes.
Description
In flight

Adults have white underparts, though the rest of the plumage goes through a complex moulting process. The male has a long pointed tail (10 to 15 cm (3.9 to 5.9 in) long) and a dark grey bill crossed by a pink band. In winter, the male has a dark cheek patch on a mainly white head and neck, a dark breast and mostly white body. In summer, the male is dark on the head, neck and back with a white cheek patch. The female has a brown back and a relatively short pointed tail. In winter, the female's head and neck are white with a dark crown. In summer, the head is dark. Juveniles resemble adult females in autumn plumage, though with a lighter, less distinct cheek patch.
Standard Measurements[12][13]
Total Body Length 440–600 mm (17.5–23.5 in)
Weight 740 g (1.63 lb)
Wingspan 710 mm (28 in)
Wing 209–228 mm (8.2–9.0 in)
Tail 165–237 mm (6.5–9.3 in)
Culmen 26–30 mm (1.0–1.2 in)
Tarsus 34–38 mm (1.3–1.5 in)

The males are vocal and have a musical yodelling call ow, ow, owal-ow.
Behaviour
Egg, Museum Wiesbaden
Mother and six ducklings in Iceland
Swimming on Lake Ontario
Breeding

Their breeding habitat is in tundra pools and marshes, but also along sea coasts and in large mountain lakes in the North Atlantic region, Alaska, northern Canada, northern Europe, and Russia. The nest is located on the ground near water; it is built using vegetation and lined with down. They are migratory and winter along the eastern and western coasts of North America, on the Great Lakes, coastal northern Europe and Asia, with stragglers to the Black Sea. The most important wintering area is the Baltic Sea, where a total of about 4.5 million gather. As of 2022 it has also been breeding in parts of Western Europe, such as on the Marker Wadden in the Netherlands.
Food and feeding

The long-tailed duck is gregarious, forming large flocks in winter and during migration. They feed by diving for mollusks, crustaceans, other marine invertebrates, freshwater insects, insect larvae, some small fish and fish eggs.[14][15] Some plant material that is also consumed includes algae, grasses, seeds and fruits in the tundra biome.[16][17] Studies show that mature adults prefer marine animals.[18][19] Specifically, they tend to eat blue mussels, baltic isopods, northern lacunas, oligochaete worms, and Amphipoda, Anostraca, Diplostraca and Mysida crustaceans, which yield higher energy per gram of live mass than other available prey.[20][21] Although they usually feed close to the surface, they are capable of diving to depths of 60 m (200 ft). According to the Audubon Society Field Guide to North American Birds they can dive to 80 fathoms (146 metres or 480 feet). They use their wings, like velvet scoters, to dive, which gives them the ability to dive much deeper than other ducks.
Status

The long-tailed duck is still hunted across a large part of its range. There has been a significant decline in the number of birds wintering in the Baltic Sea, partly due to their susceptibility to being trapped in gillnets. For these reasons the International Union for Conservation of Nature (IUCN) has categorised the long-tailed duck as vulnerable.[1] It is one of the species to which the Agreement on the Conservation of African-Eurasian Migratory Waterbirds (AEWA) applies.[22]
Notes

Though squaw originated as a word simply meaning "young woman" in the Massachusett and related Algonquian languages, it is now considered offensive by many Native Americans and is labelled as such in modern dictionaries.[9]

References

BirdLife International (2018). "Clangula hyemalis". IUCN Red List of Threatened Species. 2018: e.T22680427A132528200. doi:10.2305/IUCN.UK.2018-2.RLTS.T22680427A132528200.en. Retrieved 11 November 2021.
"coween". Merriam-Webster.com Dictionary. Merriam-Webster.
Linnaeus, Carl (1758). Systema Naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis (in Latin). Vol. 1 (10th ed.). Holmiae (Stockholm): Laurentii Salvii. p. 126.
Edwards, George (1750). A Natural History of Uncommon Birds. Vol. Part III. London: Printed for the author at the College of Physicians. p. 156 Plates 156.
Leach, William Elford (1819). Ross, John (ed.). A Voyage of Discovery made under the orders of the Admiralty in her Majesty's ships Isabella and Alexander, for the purpose of exploring Baffin's Bay, and enquiring into the probability of a North-West passage. London: John Murray. Appendix II: Zoological Memoranda, pp. 48–49.
Mayr, Ernst; Cottrell, G. William, eds. (1979). Check-List of Birds of the World. Vol. 1 (2nd ed.). Cambridge, Massachusetts: Museum of Comparative Zoology. p. 492.
Gill, Frank; Donsker, David; Rasmussen, Pamela, eds. (July 2021). "Screamers, ducks, geese & swans". IOC World Bird List Version 11.2. International Ornithologists' Union. Retrieved 1 December 2021.
Jobling, James A. (2010). The Helm Dictionary of Scientific Bird Names. London: Christopher Helm. pp. 110, 197. ISBN 978-1-4081-2501-4.
National Museum of the American Indian (2007). Do All Indians Live in Tipis?. New York: HarperCollins. p. 12. ISBN 978-0-06-115301-3.
American Ornithologists' Union (2000). "Forty-second supplement to the American Ornithologists' Union Check-list of North American Birds". The Auk. 117 (3): 847–858. doi:10.1642/0004-8038(2000)117[0847:fsstta]2.0.co;2.
Gál, Erika; Hír, János; Kessler, Eugén; Kókay, József (1998–1999). "Középsõ-miocén õsmaradványok, a Mátraszõlõs, Rákóczi-kápolna alatti útbevágásból. I. A Mátraszõlõs 1. lelõhely" [Middle Miocene fossils from the sections at the Rákóczi chapel at Mátraszőlős. Locality Mátraszõlõs I.] (PDF). Folia Historico Naturalia Musei Matraensis (in Hungarian). 23: 33–78. Archived from the original (PDF) on 2011-07-21. Retrieved 2007-02-06.
Godfrey, W. Earl (1966). The Birds of Canada. Ottawa: National Museum of Canada. p. 72.
Sibley, David Allen (2000). The Sibley Guide to Birds. New York: Knopf. p. 97. ISBN 0-679-45122-6.
Clangula hyemalis (long-tailed duck). (n.d.). Animal Diversity Web. https://animaldiversity.org/accounts/Clangula_hyemalis/
Robertson, G. J., & Savard, J. L. (2020). Long-tailed Duck (Clangula hyemalis), version 1.0. birdsoftheworld.org. https://doi.org/10.2173/bow.lotduc.01species_shared.bow.project_name
Clangula hyemalis (long-tailed duck). (n.d.). Animal Diversity Web. https://animaldiversity.org/accounts/Clangula_hyemalis/
Robertson, G. J., & Savard, J. L. (2020). Long-tailed Duck (Clangula hyemalis), version 1.0. birdsoftheworld.org. https://doi.org/10.2173/bow.lotduc.01species_shared.bow.project_name
Clangula hyemalis (long-tailed duck). (n.d.). Animal Diversity Web. https://animaldiversity.org/accounts/Clangula_hyemalis/
Robertson, G. J., & Savard, J. L. (2020). Long-tailed Duck (Clangula hyemalis), version 1.0. birdsoftheworld.org. https://doi.org/10.2173/bow.lotduc.01species_shared.bow.project_name
Clangula hyemalis (long-tailed duck). (n.d.). Animal Diversity Web. https://animaldiversity.org/accounts/Clangula_hyemalis/
Robertson, G. J., & Savard, J. L. (2020). Long-tailed Duck (Clangula hyemalis), version 1.0. birdsoftheworld.org. https://doi.org/10.2173/bow.lotduc.01species_shared.bow.project_name
"Species". Agreement on the Conservation of African-Eurasian Migratory Waterbirds (AEWA). Retrieved 14 November 2021.