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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: ParaHoxozoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Cladus: Olfactores
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Cladus: Sarcopterygii
Cladus: Rhipidistia
Cladus: Tetrapodomorpha
Cladus: Eotetrapodiformes
Cladus: Elpistostegalia
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Cladus: Sauropsida
Classis: Reptilia
Cladus: Eureptilia
Cladus: Romeriida
Cladus: Diapsida
Cladus: Neodiapsida
Cladus: Sauria
Cladus: Archelosauria
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Cladus: Crocopoda
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Cladus: Eucrocopoda
Cladus: Archosauria
Cladus: Avemetatarsalia
Cladus: Ornithodira
Cladus: Dinosauromorpha
Cladus: Dinosauriformes
Cladus: Dracohors
Cladus: Dinosauria
Cladus: Saurischia
Cladus: Theropoda
Cladus: Neotheropoda
Cladus: Averostra
Cladus: Tetanurae
Cladus: Avetheropoda
Cladus: Coelurosauria
Cladus: Tyrannoraptora
Cladus: Maniraptoromorpha
Cladus: Maniraptoriformes
Cladus: Maniraptora
Cladus: Pennaraptora
Cladus: Paraves
Cladus: Eumaniraptora
Cladus: Avialae
Subclassis: Aves
Cladus: Avebrevicauda
Cladus: Pygostylia
Cladus: Ornithothoraces
Cladus: Ornithuromorpha
Cladus: Carinatae
Cladus: Neornithes
Infraclassis: Neognathae
Cladus: Pangalloanserae
Cladus: Galloanseres
Ordo: Galliformes
Subordo: Craci

Familia: Cracidae
Subfamilia: Penelopinae
Genus: Penelope
Species: P. albipennis – P. argyrotis – P. barbata – P. bridgesi – P. dabbenei – P. jacquacu – P. jacucaca – P. marail – P. montagnii – P. obscura – P. ochrogaster – P. ortoni – P. perspicax – P. pileata – P. purpurascens – P. superciliaris

Name

Penelope Merrem, 1786

Typus: Phasianus marail Statius Müller, 1776 = Penelope marail

References
Primary references

Avium rariorum et minus cognitarum icones et descriptiones collectae e germanicus latinae factae. fasc.2 p.39

Additional references

Evangelista-Vargas, O.D. & Silveira, L.F. 2019. Morphological evidence for the taxonomic status of the Bridge’s Guan, Penelope bridgesi, with comments on the validity of P. obscura bronzina (Aves: Cracidae). Zoologia 35: 1–35. DOI: 10.3897/zoologia.35.e12993 Reference page.

Vernacular names
English: Guans
español: Pavas
suomi: Sakut
português: Jacus

Penelope is a bird genus in the family Cracidae consisting of a number of large turkey-like arboreal species, the typical guans. The range of these species is in forests from southern Mexico to tropical South America. These large birds have predominantly brown plumage and have relatively small heads when compared to the size of their bodies; they also bear a characteristic dewlap. Body lengths are typically 65 to 95 centimeters.

Most of the genus members have a typically raucous honking call. A number of the genus members are endangered species and at least one is critically endangered, usually due to tropical deforestation and hunting. In the case of several species the estimated populations are as low as a few 1000 mature birds, spread over a considerable area. Because of the scarcity of many of the genus members and also due to the habitat being often in deep or high altitude forests, little is known about some of the species habits and reproduction; in fact, some species are found at altitudes up to 3350 meters. Nests are typically built of twigs in trees.

This genus seems to have originated as part of the southward expansion of guans through the Andes and across tropical South America. Its closest relatives are probably the piping-guans, Aburria. These genera's ancestors apparently diverged some time during the Burdigalian, 20-15 Myr, but this is not corroborated by fossil evidence (Pereira et al. 2002).
Species

Image Common name Scientific Name Distribution
White-winged guan Penelope albipennis Lambayeque, Cajamarca and Piura, north-west Peru
Band-tailed guan Penelope argyrotis Colombia and Venezuela.
Bearded guan Penelope barbata Ecuador and Peru.
Yungas guan[1] Penelope bridgesi east slope of the Andes from Bolivia to northwestern Argentina
Red-faced guan Penelope dabbenei northern Argentina and in the southern border region of Bolivia.
Spix's guan Penelope jacquacu Bolivia, Brazil, Colombia, Ecuador, Guyana, Peru, and Venezuela.
White-browed guan Penelope jacucaca Caatinga in north-eastern Brazil.
Marail guan Penelope marail Brazil, French Guiana, Guyana, Suriname, and Venezuela.
Andean guan Penelope montagnii Risaralda Department of Colombia, and the Yanacocha Reserve of Ecuador.
Dusky-legged guan Penelope obscura Uruguay, northeastern Argentina and southernmost areas of Paraguay and Brazil;
Chestnut-bellied guan Penelope ochrogaster Brazil.
Baudo guan Penelope ortoni western Colombia and north-western Ecuador
Cauca guan Penelope perspicax Colombia.
White-crested guan Penelope pileata eastern Amazon basin of Brazil.
Crested guan Penelope purpurascens south Mexico and the Yucatán Peninsula to western Ecuador and southern Venezuela.
Rusty-margined guan Penelope superciliaris eastern Paraguay with extreme northeast Argentina, and eastern Bolivia in the Pantanal

References

"Species Updates – IOC World Bird List". Retrieved 2021-01-12.

Pereira, Sérgio Luiz; Baker, Allan J.& Wajntal, Anita (2002): Combined nuclear and mitochondrial DNA sequences resolve generic relationships within the Cracidae (Galliformes, Aves). Systematic Biology 51(6): 946–958. doi:10.1080/10635150290102519 PMID 12554460 PDF full text

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