Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: ParaHoxozoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Cladus: Sarcopterygii
Cladus: Rhipidistia
Cladus: Tetrapodomorpha
Cladus: Eotetrapodiformes
Cladus: Elpistostegalia
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Cladus: Synapsida
Cladus: Eupelycosauria
Cladus: Sphenacodontia
Cladus: Sphenacodontoidea
Cladus: Therapsida
Cladus: Theriodontia
Cladus: Cynodontia
Cladus: Eucynodontia
Cladus: Probainognathia
Cladus: Prozostrodontia
Cladus: Mammaliaformes
Classis: Mammalia
Cladus: Theriimorpha
Cladus: Theriiformes
Cladus: Trechnotheria
Infraclassis: †Allotheria
Ordo: †Multituberculata
Subordo: †Plagiaulacida
Familia: †Paulchoffatiidae
Subfamiliae: Paulchoffatiinae - Kuehneodontinae
Vernacular names
Paulchoffatiidae is a family of extinct mammals that lived predominantly during the Upper Jurassic period, though a couple of genera are known from the Early Cretaceous. Fossils have been reported from Europe (Portugal, Spain, Germany and England) . Paulchoffatiids were members of the order Multituberculata. They were relatively early representatives and are within the informal suborder of "Plagiaulacida". The family was named by G. Hahn in 1969, and it honors the Portuguese geologist Léon Paul Choffat. Two subfamilies are recognized.
The most productive fossil site for Paulchoffatiids has been Guimarota, Portugal. Remains from this locality are generally diagnosed on the basis of lower or upper jaws. In only one instance, that of Kuehneodon, has it been possible to match the two up. Some of the lower jaws probably represent the same animals as some of the upper, so the diversity of Paulchoffatiids is very possibly exaggerated. As the site is now a flooded, disused coalmine, further excavations are highly unlikely. However, other locations may yet provide more clarity.
The Paulchoffatiids have been further arranged into two subfamilies and a couple of other genera.
Rugosodon from the Middle Jurassic of China, known from a mostly complete skeleton, was formerly referred to the family,[1] but was later considered an indeterminate member of the Paulchoffatiid-line.[2]
Subfamily Paulchoffatiinae
Paulchoffatia, Bathmochoffatia, Guimarotodon, Henkelodon, Kielanodon, Meketibolodon, Meketichoffatia, Plesiochoffatia, Pseudobolodon, Galveodon, Sunnyodon, Xenachoffatia
Subfamily Kuehneodontinae
This taxon is restricted to one genus, Kuehneodon, with seven species.
Taxonomy
Subclass †Allotheria Marsh, 1880
Order †Multituberculata Cope, 1884:
Suborder †Plagiaulacida Simpson 1925
Family †Paulchoffatiidae Hahn, 1969
Subfamily †Paulchoffatiinae Hahn, 1971
Genus †Paulchoffatia Kühne, 1961
Species †P. delgadoi Kühne, 1961
Genus †Pseudobolodon Hahn, 1977
Species †P. oreas Hahn, 1977
Species †P. krebsi Hahn & Hahn, 1994
Genus †Henkelodon Hahn, 1987
Species †H. naias Hahn, 1987
Genus †Guimarotodon Hahn, 1969
Species †G. leiriensis Hahn, 1969
Genus †Meketibolodon (Hahn, 1978) Hahn, 1993
Species †M. robustus (Hahn, 1978) Hahn, 1993
Genus †Plesiochoffatia Hahn & Hahn, 1999
Species †P. thoas Hahn & Hahn, 1998
Species †P. peparethos Hahn & Hahn, 1998
Species †P. staphylos Hahn & Hahn, 1998
Genus †Xenachoffatia Hahn & Hahn, 1998
Species †X. oinopion Hahn & Hahn, 1998
Genus †Bathmochoffatia Hahn & Hahn, 1998
Species †B. hapax Hahn & Hahn, 1998
Genus †Kielanodon Hahn, 1987
Species †K. hopsoni Hahn, 1987
Genus †Meketichoffatia Hahn, 1993
Species †M. krausei Hahn, 1993
Genus †Galveodon Hahn & Hahn, 1992
Species †G. nannothus Hahn & Hahn, 1992
Genus †Sunnyodon Kielan-Jaworowska & Ensom, 1992
Species †S. notleyi Kielan-Jaworowska & Ensom, 1992
Subfamily †Kuehneodontinae Hahn, 1971
Genus †Kuehneodon Hahn, 1969
Species †K. dietrichi Hahn, 1969
Species †K. barcasensis Hahn & Hahn, 2001
Species †K. dryas Hahn, 1977
Species †K. guimarotensis Hahn, 1969
Species †K. hahni Antunes, 1988
Species †K. simpsoni Hahn, 1969
Species †K. uniradiculatus Hahn, 1978
References
Yuan, Chong-Xi; Ji, Qiang; Meng, Qing-Jin; Tabrum, Alan R.; Luo, Zhe-Xi (2013-08-16). "Earliest Evolution of Multituberculate Mammals Revealed by a New Jurassic Fossil". Science. 341 (6147): 779–783. doi:10.1126/science.1237970. ISSN 0036-8075. PMID 23950536. S2CID 25885140.
Martin, Thomas; O. Averianov, Alexander; A. Schultz, Julia; H. Schwermann, Achim; Wings, Oliver (2019-08-07). "Late Jurassic multituberculate mammals from Langenberg Quarry (Lower Saxony, Germany) and palaeobiogeography of European Jurassic multituberculates". Historical Biology. 33 (5): 616–629. doi:10.1080/08912963.2019.1650274. ISSN 0891-2963. S2CID 201201555.
Hahn G. and Hahn R. (2000), Multituberculates from the Guimarota mine, p. 97-107 in Martin T. and Krebs B. (eds), Guimarota - A Jurassic Ecosystem, Published by Dr Friedrich Pfeil in Münich, Germany.
Kielan-Jaworowska Z. and Hurum J.H. (2001), "Phylogeny and Systematics of multituberculate mammals". Paleontology 44, p. 389-429.
Much of this information has been derived from [1] MESOZOIC MAMMALS; Basal Multituberculata, an Internet directory.
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