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Cladus: Eukaryota
Supergroup: Opisthokonta
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Superclassis: Tetrapoda
Classis: Mammalia
Subclassis: Theria
Infraclassis: Placentalia
Ordo: Rodentia
Subordo: Myomorpha
Superfamilia: Muroidea
Familia: Muridae
Subfamilia: Murinae
Genus: Tokudaia
Species: T. muenninki - T. osimensis - T. tokunoshimensis


Tokudaia Kuroda, 1943

Type species: Rattus jerdoni osimensis Abe, 1934


* Acanthomys Tokuda, 1941
* Tokudamys Johnson, 1943


* Tokudaia on Mammal Species of the World.
Mammal Species of the World: A Taxonomic and Geographic Reference, 2 Volume Set edited by Don E. Wilson, DeeAnn M. Reeder
* Biogeographica 139: 61.

Tokudaia is a genus of murine rodent native to Japan.[1] Known as Ryūkyū spiny rats or spinous country-rats, population groups exist on several non-contiguous islands.[2] Despite differences in name and appearance, they are the closest living relatives of the Eurasian field mouse (Apodemus). Of the three species, both T. osimensis and T. tokunoshimensis have lost their Y chromosome and SRY gene; the sex chromosomes of T. muenninki, on the other hand, are abnormally large.[3]

Named species are:[3]

Muennink's spiny rat, Tokudaia muenninki
Ryukyu spiny rat, Tokudaia osimensis
Tokunoshima spiny rat, Tokudaia tokunoshimensis

At least Tokudaia osimensis may be a cryptic species complex.
See also



Musser, G.G.; Carleton, M.D. (2005). "Genus Tokudaia". In Wilson, D.E.; Reeder, D.M (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Johns Hopkins University Press. pp. 1512–1513. ISBN 978-0-8018-8221-0. OCLC 62265494.
Mammalian Genome article
Murata, C.; Yamada, F.; Kawauchi, N.; Matsuda, Y.; Kuroiwa, A. (2011-12-24). "The Y chromosome of the Okinawa spiny rat, Tokudaia muenninki, was rescued through fusion with an autosome". Chromosome Research. 20: 111–125. doi:10.1007/s10577-011-9268-6.

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