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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Cladus: Sarcopterygii
Cladus: Rhipidistia
Cladus: Tetrapodomorpha
Cladus: Eotetrapodiformes
Cladus: Elpistostegalia
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Classis: Reptilia
Cladus: Eureptilia
Cladus: Romeriida
Subclassis: Diapsida
Cladus: Sauria
Infraclassis: Lepidosauromorpha
Superordo: Lepidosauria
Ordo: Squamata
Cladus: Unidentata Episquamata Toxicofera
Subordo: Iguania
Infraordo: Acrodonta

Familia: Agamidae
Subfamilia: Draconinae
Genus: Japalura
Species (8): J. andersoniana – J. austeniana – J. dasi – J. kumaonensis – J. major – J. sagittifera – J. tricarinata – J. variegata

Japalura Gray, 1853: 387

Type species: Japalura variegata Gray, 1853, by monotypy.
Gender: feminine.

Primary references

Gray, J.E. 1853. Descriptions of some undescribed species of reptiles collected by Dr. Joseph Hooker in the Khassia Mountains, East Bengal, and Sikkim Himalaya. Annals and Magazine of Natural History. Series 2 12(72): 386–392. BHL Reference page.

Additional references

Denzer, W., Manthey, U. & Campbell, P.D. 2019. Catalogue of type specimens of the agamid lizard genus Japalura s. l. (Squamata: Agamidae: Draconinae). Zootaxa 4612(1): 109–125. DOI: 10.11646/zootaxa.4612.1.8. Reference page.
Wang, K., Che, J., Lin, S.-M., Deepak, V., Datta-Roy, A., Jiang, K., Jin, J.-Q., Chen, H.-M. & Siler, C.D. 2019. Multilocus phylogeny and revised classification for mountain dragons of the genus Japalura s.l. (Reptilia: Agamidae: Draconinae) from Asia. Zoological Journal of the Linnean Society 185(1): 246–267. DOI: 10.1093/zoolinnean/zly034 Open access Reference page.
Gowande, G.G., Bhosale, H.S., Phansalkar, P.U., Sawant, M. & Mirza, Z.A. 2021. On the systematics and the phylogenetic position of the poorly known, montane dragon-lizard species Pseudocalotes austeniana (Annandale, 1908) (Squamata, Agamidae, Draconinae). Evolutionary Systematics 5(1): 141–150. DOI: 10.3897/evolsyst.5.67137 Open access Reference page.


Uetz, P. & Hallermann, J. 2022. Japalura . The Reptile Database. Accessed on 2 November 2019.

Vernacular names
English: Himalayan Dragons

Japalura is a genus of lizards in the family Agamidae. Species of Japalura are native to Pakistan, India, China, and Myanmar.[1][2] Many species have been moved to the genus Diploderma.

The following eight species are recognized as being valid:[3]

Japalura andersoniana Annandale, 1905 – Anderson's mountain lizard
Japalura austeniana (Annandale, 1908) – Abor Hills agama
Japalura dasi (Shah & Kästle, 2002) – Agaupani mountain lizard, Agaupani forest agama
Japalura kumaonensis (Annandale, 1907) – Kumaon mountain lizard, Kumaon forest agama
Japalura major (Jerdon, 1870) – large mountain lizard, greater forest agama
Japalura sagittifera M.A. Smith, 1940 – Burmese japalure
Japalura tricarinata (Blyth, 1853) – three-keeled mountain lizard, tricarinate forest agama
Japalura variegata Gray, 1853 – variegated mountain lizard, Himalayan dragon

Nota bene: a binomial authority in parentheses indicates that the species was originally described in a genus other than Japalura.

Kunte K, Manthey U (2009). "Rediscovery of Japalura sagittifera (Sauria: Agamidae) from the eastern Himalayas, Arunachal Pradesh: An addition to the Indian herpetofauna" (PDF). Sauria. 31 (2): 49–55.
Mahony S (2009). "A new species of Japalura (Reptilia: Agamidae) from northeast India with a discussion of the similar species Japalura sagittifera Smith, 1940 and Japalura planidorsata Jerdon, 1870" (PDF). Zootaxa. 2212: 41–61. doi:10.11646/zootaxa.2212.1.2.

Japalura at the Reptile Database. Accessed 18 October 2018.

Further reading

Gray JE (1853). "Descriptions of some undescribed species of Reptiles collected by Dr. Joseph Hooker in the Khassia Mountains, East Bengal, and Sikkim Himalaya". Annals and Magazine of Natural History, Second Series 12: 386-392. (Japalura, new genus, pp. 387–388).

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