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Empidoidea

Cladus: Eukaryota
Supergroup: Opisthokonta
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Cladus: Pterygota
Cladus: Metapterygota
Cladus: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Panorpida
Ordo: Diptera
Subordo: Brachycera
Infraordo: Asilomorpha
Superfamilia: Empidoidea
Familiae: Atelestidae - Brachystomatidae - Dolichopodidae - Empididae - Hybotidae

Name

Empidoidea

References

* Sinclair, B.J.; Cumming, J.M. 2006: The morphology, higher-level phylogeny and classification of the Empidoidea (Diptera). Zootaxa, 1180: 1-172. [1] [2]

Vernacular names
日本語: オドリバエ上科

The Empidoidea are a large monophyletic superfamily of true flies, the sister taxon to the Muscomorpha (= "Cyclorrhapha"). These two groups are sometimes united in the unranked taxon Eremoneura. There are some 10,000 known species in the Empidoidea[1].

The majority of these insects are predatory, often with large compound eyes (sometimes covering almost the entire surface of the head), and tend to be associated with moist, temperate habitats. Many have a well-developed anal lobe in the wings, and/or a small but distinct anal cell.

Systematics

The most familiar families in the group are the Empididae and Dolichopodidae, both of which occur worldwide and contain thousands of species. The smaller families were formerly included in the Empididae, but have since been accorded family status[2]. The Brachystomatinae and Microphorinae are sometimes elevated to full-blown families too, but this would apparently make the Empididae and Dolichopodidae paraphyletic and thus they are retained as subfamilies here.

It seems that the present systematic treatment essentially reflects phylogeny rather well; there are some of the more basal taxa in each lineage which are of more uncertain position in cladistic analyses, but at least as an interim solution, the four-family approach seems most warranted. Arranged in phylogenetic sequence, the families are:[1]

* Atelestidae
* Hybotidae – dance flies
* Dolichopodidae – long-legged flies (including Microphoridae)
* Empididae – dagger flies and balloon flies (including Brachystomatidae)

Footnotes

1. ^ a b Moulton & Wiegmann (2007)
2. ^ Sinclair & Cumming (2006)

References

* Moulton, J.K. & Wiegmann, B.M. (2007): The phylogenetic relationships of flies in the superfamily Empidoidea (Insecta: Diptera). Mol. Phylogenet. Evol. 43(3): 701-713. doi:10.1016/j.ympev.2007.02.029 (HTML abstract)
* Sinclair, B.J. & Cumming, J.M. (2006): The morphology, higher-level phylogeny and classification of the Empidoidea (Diptera). Zootaxa 1180: 1-172. PDF fulltext

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