Psammodius

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Coleopterida
Ordo: Coleoptera
Subordo: Polyphaga
Infraordo: Scarabaeiformia
Superfamilia: Scarabaeoidea

Familia: Scarabaeidae
Subfamilia: Aphodiinae
Tribus: Psammodiini
Subtribus: Psammodiina
Genus: Psammodius
Species (27): P. asper – P. asperoides – P. basalis – P. besucheti – P. bilyi – P. bulgaricus – P. caucasicus – P. convexus – P. danubialis – P. flavolittoralis – P. generosus – P. hangangnensis – P. kobayashii – P. laevipennis – P. maceki – P. macnamarae – P. maruyamai – P. nepalensis – P. nocturnus – P. ovatus – P. pierottii – P. plicicollis – P. schmidti – P. sefrensis – P. seiferti – P. sungshinarum – P. tesari
Name

Psammodius Fallén, 1807

Type species:

References
Additional references

Ochi, T.; Kawahara, M.; Inagaki, M. 2011: Taxonomic notes on some Japanese coprophagous lamellicorn beetles (Coleoptera, Scarabaeoidea) (8) Descriptions of a new species of the genus Psammodius and a new species of the genus Trichiorhyssemus (Aphodiidae) from Tanegashima Is., Southwest Japan. Kogane, (12 ): 69–77. [not seen]
Pittino, R. 2006: Redescription of the genus Petrovitzius Rakovič, 1979, and descriptions of Rakovicius gen. nov. from East Asia, and Rakovicius kawaii sp. nov. from Taiwan (Coleoptera, Aphodiidae, Psammodiinae). Kogane, (7 ): 11–21. [Publication date: 'Jul. 2006', sourced from first page of article]
Pittino, R. 2007: A review of the Western Palaearctic species of the genus Psammodius Fallén, 1807, with description of a new species from Eastern Greece (Coleoptera Aphodiidae Psammodiinae). Giornale italiano di entomologia, 12: 93–117. [not seen]
Pittino, R.; Shokhin I.V. 2006: A new species of the genus Psammodius Fallén, 1807 from Northeastern Anatolia and Caucasus. (Coleoptera, Aphodiidae, Psammodiinae). Kogane, (7 ): 23–26.

Links

Schoolmeesters, P. 2017. Scarabs: World Scarabaeidae Database (version Jul 2016). In: Roskov Y., Abucay L., Orrell T., Nicolson D., Bailly N., Kirk P., Bourgoin T., DeWalt R.E., Decock W., De Wever A., Nieukerken E. van, Zarucchi J., Penev L., eds. 2017. Species 2000 & ITIS Catalogue of Life, 30th January 2017. Digital resource at www.catalogueoflife.org/col. Species 2000: Naturalis, Leiden, the Netherlands. ISSN 2405-8858. Reference page. [accessed on April 18, 2017]

Psammodius is a genus of aphodiine dung beetles in the family Scarabaeidae. There are at least 3 described species in Psammodius.[1][2]
Species

Psammodius basalis (Mulsant & Rey, 1870)
Psammodius laevipennis Costa, 1844
Psammodius pierottii Pittino, 1979

References

"Psammodius Genus Information". BugGuide.net. Retrieved 1 February 2018.

"Psammodius Report". Integrated Taxonomic Information System. Retrieved 1 February 2018.

Further reading

Arnett, R.H. Jr., M. C. Thomas, P. E. Skelley and J. H. Frank. (eds.). (2002). American Beetles, Volume II: Polyphaga: Scarabaeoidea through Curculionoidea. CRC Press LLC, Boca Raton, FL.
Ross H. Arnett (30 July 2000). American Insects: A Handbook of the Insects of America North of Mexico. CRC Press. ISBN 978-0-8493-0212-1.
Richard E. White. (1983). Peterson Field Guides: Beetles. Houghton Mifflin Company.