Zygadenia

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Coleopterida
Ordo: Coleoptera
Subordo: Archostemata
Superfamilia: Cupedoidea

Familia: Ommatidae
Subfamilia: Ommatinae
Tribus: †Notocupedini
Genus: Zygadenia
Species: Z. floodpagei – Z. homora – Z. liui – Z. martinae – Z. simpsoni – Z. westraliensis – ...
Name

†Zygadenia Handlirsch, 1906
Synonyms

†Kakoselia Handlirsch, 1906 [syn. Ponomarenko (1995: 95)]
†Sinocupes Lin, 1976 [syn. Ponomarenko (1995: 95)]
†Conexicoxa Lin, 1986 [syn. Ponomarenko (1995: 95)]
†Forticupes Hong & Wang, 1990 [syn. Ponomarenko (1995: 95)]
†Picticupes Hong & Wang, 1990 [syn. Ponomarenko (1995: 95)]
†Lupicupes Ren, 1995 [syn. Ponomarenko (1995: 95)]

References

Jarzembowski, E.A. et al. 2015 [2014 epub]. Boring beetles are not necessarily dull: New notocupedins (Insecta: Coleoptera) from the Mesozoic of Eurasia and East Gondwana. Cretaceous research 52(B): 431–439. DOI: 10.1016/j.cretres.2014.03.006 Paywall. Reference page.
Kirejtshuk, A.G., Ponomarenko, A.G., Prokin, A.A., Chang, H., Nikolajev, G.V. & Ren, D. 2010. Current knowledge of Mesozoic Coleoptera from Daohugou and Liaoning (northeast China). Acta Geologica Sinica - English Edition 84(4): 783–792. DOI: 10.1111/j.1755-6724.2010.00253.x Paywall. Reference page.
Martin, S.K. 2010: Early Jurassic coleopterans from the Mintaja insect locality, Western Australia. Acta geologica sinica - English edition, 84 (4): 925–953. DOI: 10.1111/j.1755-6724.2010.00276.x
Ponomarenko, A.G. 2006: On the types of Mesozoic archostematan beetles (Insecta, Coleoptera, Archostemata) in the Natural History Museum, London. Paleontological journal, 40: 90–99.
Ponomarenko, A.G.; Ren, D. 2010: First record of Notocupes (Coleoptera: Cupedidae) in locality Daohugou, Middle Jurassic of Inner Mongolia, China. Annales zoologici, 60: 169–171. ISSN: 0003-4541 DOI: 10.3161/000345410X516812
Ren, D. 1995: Fossil insects. In Faunae and stratigraphy of the Jurassic-Cretaceous in Beijing and adjacent areas, Beijing. Seismic, Beijing.
Wang, W.-l.; Liu, M.-w. 1996: A new species of Notocupes from the Cretaceous of Laiyang Basin, Shandong Province. Memoirs of Beijing Natural History Museum, (55 ): 79–82.

Links

ION search for Notocupes

Zygadenia is an extinct genus of beetles in the family Ommatidae.

Triassic Species

After Kirejtshuk, 2020. Kirejtshuk notes that the two Lower Jurassic Australian species Z. westraliensis and Z. martinae possibly merit placement in their own separate genus.[1]
Ladinian (242 Ma to ~237 Ma)

Tongchuan Formation, China

Z. shiluoensis Hong 1984

Carnian (237 Ma to 227 Ma)

Koldzat Formation, Kazakhstan

Z. oxypygus (Ponomarenko, 1969)

Madygen Formation, Kyrgyzstan

Z. laticella (Ponomarenko, 1969)
Z. rostratus (Ponomarenko, 1969)
Z. tenuis (Ponomarenko, 1969)

Jurassic species
Hettangian (201.3 Ma to 199.3 Ma)

Dzhil Formation, Kyrgyzstan

Z. cellulosa (Ponomarenko, 1969)
Z. issykkulensis (Ponomarenko, 1969)
Z. kirghizicus (Ponomarenko, 1969)
Z. latus (Ponomarenko, 1969)
Z. sogutensis (Ponomarenko, 1969)

Guanyintan Formation, China

Z. homorus (Lin, 1986)

Toarcian (182.7 Ma to 174.1 Ma)

Cattamarra Coal Measures, Australia

Z. martinae Jarzembowski et al. 2015
Z. westraliensis Riek 1968

Guanyintan Formation, China

Z. lini (Ponomarenko et al. 2012)

Sulyukta Formation, Kyrgyzstan

Z. crassus (Ponomarenko, 1969)

Bajocian (170.3 Ma to 168.3 Ma)

Bakhar Formation, Mongolia

Z. brachycephalus (Ponomarenko, 1994)
Z. exiguus (Ponomarenko, 1994)
Z. longicollis (Ponomarenko, 1994)

Callovian (166.1 Ma to 163.5 Ma)

Haifanggou Formation, China

Z. dischides (Zhang 1986)
Z. jurassica Hong 1983
Z. pingi (Ponomarenko and Ren 2010)

Tyumen Formation, Russian

Z. patulus Ponomarenko, 1985

Oxfordian (163.5 Ma to 157.3 Ma)

Karabastau Formation, Kazakhstan

Z. foersteri (Ponomarenko, 1971)
Z. lapidarius (Ponomarenko, 1968)
Z. nigromonticola (Ponomarenko, 1968)
Z. picturatus (Ponomarenko, 1964)
Z. pulcher (Ponomarenko, 1968)

Tithonian (152.1 Ma To 145.0 Ma)

Shar Teeg, Mongolia

Z. brachycephalus (Ponomarenko, 1994)
Z. exiguus (Ponomarenko, 1994)
Z. giebeli Ponomarenko 2014
Z. handlirschi Ponomarenko 2014

Solnhofen, Germany

Z. reticulatus (Oppenheim, 1888)
Z. tripartitus (Oppenheim, 1888)

Cretaceous species
Berriasian (145.0 Ma To 139.8 Ma)

Lulworth Formation, United Kingdom

Z. angliae (Giebel 1856)
Z. tuberculata (Giebel, 1856)

Hauterivian (132.9 Ma to 129.4 Ma)

Dabeigou Formation, China

Z. lentus (Ren, 1995)

Weald Clay, United Kingdom

Z. angliae (Giebel 1856)
Z. floodpagei Jarzembowski et al. 2015
Z. simpsoni Jarzembowski et al. 2015

Barremian (129.4 Ma to 125.0 Ma)

Jianshangou Formation, China

Z. laetus (Lin, 1976)

La Huérguina Formation, Las Hoyas, Spain

Z. diazromerali Soriano & Delclòs, 2006
Z. longicoxa Soriano & Delclòs, 2006
Z. siniestri Soriano & Delclòs, 2006
Z. viridis Soriano & Delclòs, 2006

La Pedrera de Rúbies Formation, Las Hoyas, Spain

Z. martinclosas Soriano & Delclòs, 2006
Z. oculata Soriano & Delclòs, 2006

Ulan-Argalant Formation, Mongolia

Z. mongolicus (Ponomarenko, 1994)

Weald Clay, United Kingdom

Z. angliae (Giebel 1856)
Z. floodpagei Jarzembowski et al. 2015
Z. tuberculata (Giebel 1856)
Z. simpsoni Jarzembowski et al. 2015

Aptian (125.0 to ~113.0 Ma)

Argun Formation, Russia

Z. semen (Ponomarenko, 2000)
Z. sibirica Ponomarenko, 2000
Z. sinitzae Ponomarenko, 2000

Dzun-Bain Formation, Mongolia

Z. dundulaensis (Ponomarenko, 1994)
Z. elegans (Ponomarenko, 1994)

Jehol Group, China

Z. valida Lin 1976

Khasurty Formation, Russia

Z. khasurtyiensis (Strelnikova 2019)

Laiyang Formation, China

Z. laiyangensis (Hong, 1990)
Z. ludongensis (Wang and Liu 1996)
Z. tuanwangensis (Hong, 1990)

Shahai Formation, China

Z. kezuoensis Hong 1987

Yixian Formation, China

Z. aliena (Tan and Ren 2006)
Z. epicharis (Tan et al. 2005)
Z. eumeura (Tan et al. 2006)
Z. liui Jarzembowski et al. 2015
Z. miniscula Tan et al. 2006
Z. porrecta Tan et al. 2006
Z. protensa Tan et al. 2006
Z. psilata Tan et al. 2005
Z. rudis Tan et al. 2005
Z. stabilis Tan et al. 2005

Zaza Formation, Russia

Z. caudatus (Ponomarenko, 1966)
Z. excellens (Ponomarenko, 1966)
Z. vitimensis (Ponomarenko, 1966)

Turonian (93.9 to 89.8 Ma)

Emanra Formation, Russia

Z. khetanensis (Ponomarenko in Cromov et al., 1993)

Kzyl-Zhar Locality, Kazakhstan

Z. caduca (Ponomarenko, 1969)

References

Kirejtshuk, Alexander G. (2020). "Taxonomic Review of Fossil Coleopterous Families (Insecta, Coleoptera). Suborder Archostemata: Superfamilies Coleopseoidea and Cupedoidea". Geosciences. 10 (2): 73. Bibcode:2020Geosc..10...73K. doi:10.3390/geosciences10020073. ISSN 2076-3263.