Coelotes

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Chelicerata
Classis: Arachnida
Ordo: Araneae
Subordo: Opisthothelae
Infraordo: Araneomorphae
Taxon: Neocribellatae
Series: Entelegynae
Superfamilia: Agelenoidea

Familia: Agelenidae
Subfamilia: Coelotinae
Genus: Coelotes
List of species
A

C. acerbus – C. adnexus – C. aguniensis – C. akakinaensis – C. albimontanus – C. alpinus – C. amamiensis – C. amplilamnis – C. antri – C. arganoi – C. aritai – C. atropos –
B

C. bifida – C. bifurcatus – C. biprocessis – C. brachiatus – C. brevis –
C

C. capacilimbus – C. caudatus – C. cavernalis – C. cavicola – C. charitonovi – C. chenzhou – C. chishuiensis – C. cochleariformis – C. coenobita – C. colosseus – C. conversus – C. cornutus – C. cristiformis – C. curvilamnis – C. cylistus –
D

C. decolor – C. degeneratus – C. doii – C. dormans –
E

C. eharai – C. enasanus – C. everesti – C. exaptus – C. exilis – C. exitialis –
F

C. fanjingensis – C. filamentaceus – C. forficatus – C. fujian – C. furvus –
G

C. galeiformis – C. gifuensis – C. globasus – C. gotoensis – C. guangxian – C. guizhouensis – C. guttatus –
H

C. hachijoensis – C. hamamurai – C. hataensis – C. hengshanensis – C. hexommatus – C. hikonensis – C. hiradoensis – C. hiratsukai – C. hiruzenensis – C. hiurai –
I

C. ibukiensis – C. icohamatus – C. iharai – C. iheyaensis – C. ikiensis – C. improprius – C. inabaensis – C. indentatus – C. indistinctus – C. insulanus – C. introhamatus – C. iriei – C. isensis – C. italicus – C. iyoensis – C. izenaensis –
J

C. jianfenglingensis – C. jucundus – C. juglandicola –
K

C. kagaensis – C. kakeromaensis – C. katsurai – C. keramaensis – C. kimi – C. kintaroi – C. kirgisicus – C. kitazawai – C. koshikiensis – C. kumejimanus – C. kumensis – C. kunigamiensis –
L

C. lamellatus – C. laohuanglongensis – C. latus – C. ledongensis – C. lini – C. longus – C. luculli –
M

C. maculatus – C. major – C. mastrucatus – C. mediocris – C. micado – C. microps – C. minobusanus – C. minoensis – C. miyakoensis – C. modestus – C. mohrii – C. motobuensis – C. multannulatus – C. musashiensis –
N

C. nagaraensis – C. nasensis – C. nazuna – C. nenilini – C. ningmingensis – C. noctulus – C. notoensis –
O

C. obako – C. obesus – C. obtusangulus – C. ogatai – C. okinawensis – C. osamui – C. osellai – C. oshimaensis – C. oxyacanthus –
P

C. pabulator – C. parallelus – C. pastoralis – C. pastor – C. pedodentalis – C. perbrevis – C. personatus – C. pervicax – C. phthisicus – C. pickardi – C. plancyi – C. poleneci – C. polyedricus – C. poricus – C. porus – C. poweri – C. processus – C. progressoridentes – C. prolixus –
Q

C. quadratus –
R

C. rhododendri – C. robustior – C. robustus – C. rudolfi – C. rugosus –
S

C. saccatus – C. saikaiensis – C. samaksanensis – C. sanoi – C. satoi – C. sawadai – C. semicircularis – C. septus – C. serpentinus – C. shimajiriensis – C. simoni – C. simplex – C. sinopensis – C. sinuolatus – C. solitarius – C. songae – C. sordidus – C. striatilamnis – C. stylifer – C. suruga – C. suthepicus –
T

C. takanawaensis – C. tarumii – C. taurus – C. tegenarioides – C. tenutubilaris – C. terrestris – C. tiantangensis – C. tiantongensis – C. titaniacus – C. tochigiensis – C. tojoi – C. tokaraensis – C. tokunoshimaensis – C. tominagai – C. tonakiensis – C. transiliensis – C. troglocaecus – C. tumidivulvus – C. turkestanicus –
U

C. uncatus – C. undulatus – C. unicatus – C. unzenensis – C. uozumii –
V

C. vallei – C. vestigialis – C. vignai –
W

C. wangi – C. wugeshanensis –
X

C. xinjiangensis –
Y/Z

C. yaginumai – C. yahagiensis – C. yambaruensis – C. yanlingensis – C. yodoensis – C. yunnanensis – C. zaoensis
Nomina dubia

Coelotes ghigii Caporiacco, 1948d: 235 (j, Italy) -- de Blauwe, 1973a: 39.
Coelotes immaculatus O. Pickard-Cambridge, 1873b: 444 (f, Britain) -- Bristowe, 1941: 518.
Coelotes pastor pirinicus Drensky, 1942: 43, 59, f. 9c (f, Bulgaria) -- Deltshev & Blagoev, 2001: 111, after Deltshev, 1990: 42.

Name

Coelotes Blackwall, 1841

Type species: Drassus atropos Charles Athanase Walckenaer, 1830
Synonyms

Urobia Komatsu, 1957

Type species: Urobia hexommata Toshihiro Komatsu, 1957

References
Primary references

Blackwall, J. 1841: Trans. Linn. Soc. Lond. 18: 601–670. [618]

Additional references

Liu, J. & S.Q. Li, 2009: New Cave-Dwelling Coelotine Spiders from the Yunnan-Guizhou Plateau, China (Araneae: Amaurobiidae). Zoological Studies 48 (5): 665–681. Full article: [1].
Liu, J.; Li, S. 2010: New coelotine spiders from Xishuangbanna Rainforest, southwestern China (Araneae: Amaurobiidae). Zootaxa, 2442: 1–24. Preview
Liu, J.H., Xiao, Y.H., Zhang, M.Z., Xu, X. & Liu, K.K. 2021. Four new coelotine species (Araneae, Agelenidae, Coelotinae) from South China, with the first description of the male of Coelotes septus Wang, Yin, Peng & Xie, 1990. ZooKeys 1029: 93–112. DOI: 10.3897/zookeys.1029.63060. Open access. Reference page.
Luo, Y. & Chen, J. 2015. Two new Coelotes species from China (Araneae: Agelenidae: Coelotinae). Zoological Systematics 40(3): 258–267. DOI: 10.11865/zs.20150303. Reference page.
Nishikawa, Y. 2009. A new genus and 44 new species of the family Coelotidae (Arachnidae, Araneae) from Japan. In: Ono, H. (ed.) The spiders of Japan with keys to the families and genera and illustrations of the species. Tokai University Press, Kanagawa: 51-70. Reference page.
Okumura, K-i. 2010: First description of the male of Platocoelotes uenoi (Yamaguchi & Yaginuma 1971), n. comb. and a new record of Draconarius coreanus (Paik & Yaginuma 1969) (Araneae: Coelotidae) from Kyushu, Japan. Acta arachnologica, 59(1): 5–7. DOI: 10.2476/asjaa.59.5
Okumura, K.I., Suzuki, Y. & Serita, R. 2020. Two new species of spiders of the families Agelenidae and Amaurobiidae from Okinawa Island, Japan. Arachnology 18(6): 596-601. DOI: 10.13156/arac.2020.18.6.596. Open access. Reference page.
Shimojana, M., Nishihira, M. 2000: A New Cave-Dwelling Eyeless Spider of the Genus Coelotes (Araneae: Amaurobiidae) from Okinawa Island, the Ryukyu Islands, Japan, with Notes on Possible Parthenogenesis. Acta Arachnologica 49 (1): 29-40. full article (pdf). Reference page.
Wang, X.-p.; Griswold, C.E.; Ubick, D. 2009: On the pseudoterrestris species group of the spider genus Coelotes (Araneae, Amaurobiidae). Zootaxa, 2313: 1–34. Abstract & excerpt
Wang, X.-P. ; I.-M. Tso & H.-Y. Wu, 2001: Three new Coelotes spiders (Araneae: Amaurobiidae) from Taiwan. Zoological Studies 40 (2): 127–133. Full article: [2].
Zhu, C.D. & Wang, J.F. 1994. Seven new species of the genus Coelotes from China (Araneae: Agelenidae). Acta Zootaxonomica Sinica 19: 37-45. Reference page.

Links

Platnick, N. I. 2009. The World Spider Catalog, version 9.5. American Museum of Natural History. [3]

Vernacular names
한국어: 기슭비탈거미속

Coelotes is a genus of funnel weavers first described by John Blackwall in 1841.[4] A large number of species are found throughout Europe and Asia.[1]
Species

As of April 2019 it contains 189 species:[1]

C. acerbus Liu, Li & Pham, 2010 – Vietnam
C. adnexus Zhang, Zhu & Wang, 2017 – China
C. aguniensis Shimojana, 2000 – Japan (Ryukyu Is.)
C. akakinaensis Shimojana, 2000 – Japan (Ryukyu Is.)
C. albimontanus Nishikawa, 2009 – Japan
C. alpinus Polenec, 1972 – Italy, Austria, Slovenia
C. amamiensis Shimojana, 1989 – Japan (Ryukyu Is.)
C. amplilamnis Saito, 1936 – China
C. antri (Komatsu, 1961) – Japan
C. arganoi Brignoli, 1978 – Turkey
C. aritai Nishikawa, 2009 – Japan
C. atropos (Walckenaer, 1830) – Europe
C. bifurcatus Okumura & Ono, 2006 – Japan
C. biprocessis Zhang, Zhu & Wang, 2017 – China
C. brachiatus Wang, Yin, Peng & Xie, 1990 – China
C. brevis Xu & Li, 2007 – China
C. capacilimbus Xu & Li, 2006 – China
C. caudatus de Blauwe, 1973 – Lebanon
C. cavicola (Komatsu, 1961) – Japan
C. charitonovi Spassky, 1939 – Central Asia
C. chenzhou Zhang & Yin, 2001 – China
C. chishuiensis Zhang, Zhu & Wang, 2017 – China
C. coenobita Brignoli, 1978 – Turkey
C. colosseus Xu & Li, 2007 – China
C. conversus Xu & Li, 2006 – China
C. cornutus Nishikawa, 2009 – Japan
C. cristiformis Jiang, Chen & Zhang, 2018 – China
C. curvilamnis Ovtchinnikov, 2000 – Kyrgyzstan
Coelotes c. alatauensis Ovtchinnikov, 2000 – Kazakhstan, Kyrgyzstan
Coelotes c. boomensis Ovtchinnikov, 2000 – Kyrgyzstan
C. cylistus Peng & Wang, 1997 – China
C. decolor Nishikawa, 1973 – Japan
C. degeneratus Liu & Li, 2009 – China
C. doii Nishikawa, 2009 – Japan
C. dormans Nishikawa, 2009 – Japan
C. eharai Arita, 1976 – Japan
C. enasanus Nishikawa, 2009 – Japan
C. everesti Hu, 2001 – China
C. exaptus Banks, 1898 – Mexico
C. exilis Nishikawa, 2009 – Japan
C. exitialis L. Koch, 1878 – Korea, Japan
C. fanjingensis Zhang, Zhu & Wang, 2017 – China
C. fujian Zhang, Zhu & Wang, 2017 – China
C. furvus Liu, Li & Pham, 2010 – Vietnam
C. galeiformis Wang, Yin, Peng & Xie, 1990 – China
C. gifuensis Nishikawa, 2009 – Japan
C. globasus (Wang, Peng & Kim, 1996) – China
C. gotoensis Okumura, 2007 – Japan
C. guttatus Wang, Yin, Peng & Xie, 1990 – China
C. hachijoensis Ono, 2008 – Japan
C. hamamurai Yaginuma, 1967 – Japan
C. hataensis Nishikawa, 2009 – Japan
C. hexommatus (Komatsu, 1957) – Japan
C. hikonensis Nishikawa, 2009 – Japan
C. hiradoensis Okumura & Ono, 2006 – Japan
C. hiratsukai Arita, 1976 – Japan
C. hiruzenensis Nishikawa, 2009 – Japan
C. hiurai Nishikawa, 2009 – Japan
C. ibukiensis Nishikawa, 2009 – Japan
C. icohamatus Zhu & Wang, 1991 – China
C. iharai Okumura, 2007 – Japan
C. iheyaensis Shimojana, 2000 – Japan (Ryukyu Is.)
C. ikiensis Nishikawa, 2009 – Japan
C. improprius (Wang, Griswold & Miller, 2010) – China
C. indentatus Zhang, Zhu & Wang, 2017 – China
C. insulanus Shimojana, 2000 – Japan (Ryukyu Is.)
C. introhamatus Xu & Li, 2006 – China
C. iriei Okumura, 2013 – Japan
C. italicus Kritscher, 1956 – Italy
C. iyoensis Nishikawa, 2009 – Japan
C. izenaensis Shimojana, 2000 – Japan (Ryukyu Is.)
C. jianfenglingensis (Liu & Li, 2009) – China
C. juglandicola Ovtchinnikov, 1984 – Kyrgyzstan
C. kagaensis Nishikawa, 2009 – Japan
C. kakeromaensis Shimojana, 2000 – Japan (Ryukyu Is.)
C. katsurai Nishikawa, 2009 – Japan
C. keramaensis Shimojana, 2000 – Japan (Ryukyu Is.)
C. kimi Kim & Park, 2009 – Korea
C. kintaroi Nishikawa, 1983 – Japan
C. kirgisicus Ovtchinnikov, 2000 – Kyrgyzstan
C. kitazawai Yaginuma, 1972 – Japan
C. koshikiensis Okumura, 2013 – Japan
C. kumejimanus Shimojana, 2000 – Japan (Ryukyu Is.)
C. kumensis Shimojana, 1989 – Japan (Ryukyu Is.)
C. lamellatus Nishikawa, 2009 – Japan
C. laohuanglongensis Liu & Li, 2009 – China
C. ledongensis Zhang, Zhu & Wang, 2017 – China
C. luculli Brignoli, 1978 – Turkey
C. maculatus Zhang, Peng & Kim, 1997 – China
C. mastrucatus Wang, Yin, Peng & Xie, 1990 – China
C. mediocris Kulczyński, 1887 – Switzerland, Italy, Ukraine?
C. micado Strand, 1907 – Japan
C. microps Schenkel, 1963 – China
C. minobusanus Nishikawa, 2009 – Japan
C. minoensis Nishikawa, 2009 – Japan
C. miyakoensis Shimojana, 2000 – Japan (Ryukyu Is.)
C. modestus Simon, 1880 – China, Japan
C. mohrii Nishikawa, 2009 – Japan
C. motobuensis Shimojana, 2000 – Japan (Ryukyu Is.)
C. multannulatus Zhang, Zhu, Sun & Song, 2006 – China
C. musashiensis Nishikawa, 1989 – Japan
C. nagaraensis Nishikawa, 2009 – Japan
C. nasensis Shimojana, 2000 – Japan (Ryukyu Is.)
C. nazuna Nishikawa, 2009 – Japan
C. nenilini Ovtchinnikov, 1999 – Uzbekistan
C. ningmingensis Peng, Yan, Liu & Kim, 1998 – China
C. noctulus Wang, Yin, Peng & Xie, 1990 – China
C. notoensis Nishikawa, 2009 – Japan
C. obako Nishikawa, 1983 – Japan
C. obtusangulus Luo & Chen, 2015 – China
C. ogatai Nishikawa, 2009 – Japan
C. okinawensis Shimojana, 1989 – Japan (Ryukyu Is.)
C. osamui Nishikawa, 2009 – Japan
C. osellai de Blauwe, 1973 – Italy
C. oshimaensis Shimojana, 2000 – Japan (Ryukyu Is.)
C. oxyacanthus Okumura, 2013 – Japan
C. pabulator Simon, 1875 – France, Switzerland
C. pastoralis Ovtchinnikov, 2000 – Kazakhstan, Kyrgyzstan
C. pedodentalis Zhang, Zhu, Sun & Song, 2006 – China
C. perbrevis Liu, Li & Pham, 2010 – Vietnam
C. personatus Nishikawa, 1973 – Japan
C. pervicax Hu & Li, 1987 – China
C. phthisicus Brignoli, 1978 – Turkey
C. pickardi O. Pickard-Cambridge, 1873 – Switzerland, Italy
Coelotes p. carpathensis Ovtchinnikov, 1999 – Ukraine
Coelotes p. pastor Simon, 1875 – France
Coelotes p. tirolensis (Kulczyński, 1906) – Switzerland, Italy, Ukraine?
C. poleneci Wiehle, 1964 – Austria, Slovenia
C. polyedricus Liu, Li & Pham, 2010 – Vietnam
C. poricus Zhang, Zhu & Wang, 2017 – China
C. poweri Simon, 1875 – France
C. processus Xu & Li, 2007 – China
C. progressoridentes Ovtchinnikov, 2000 – Kyrgyzstan
C. quadratus Wang, Yin, Peng & Xie, 1990 – China
C. rhododendri Brignoli, 1978 – Turkey
C. robustior Nishikawa, 2009 – Japan
C. robustus Wang, Yin, Peng & Xie, 1990 – China
C. rudolfi (Schenkel, 1925) – Switzerland
C. rugosus (Wang, Peng & Kim, 1996) – China
C. saccatus Peng & Yin, 1998 – China
C. saikaiensis Okumura, 2013 – Japan
C. sanoi Nishikawa, 2009 – Japan
C. sawadai Nishikawa, 2009 – Japan
C. septus Wang, Yin, Peng & Xie, 1990 – China
C. serpentinus Jiang, Chen & Zhang, 2018 – China
C. shimajiriensis Shimojana, 2000 – Japan (Ryukyu Is.)
C. simplex O. Pickard-Cambridge, 1885 – China (Yarkand)
C. sinopensis Danişman, Karanfil & Coşar, 2016 – Turkey
C. sinuolatus Zhang, Zhu & Wang, 2017 – China
C. solitarius L. Koch, 1868 – Europe
C. songae Liu, Li & Pham, 2010 – Vietnam
C. sordidus Ovtchinnikov, 2000 – Kazakhstan, Kyrgyzstan
C. striatilamnis Ovtchinnikov, 2000 – Kazakhstan, Kyrgyzstan
Coelotes s. ketmenensis Ovtchinnikov, 2001 – Kazakhstan
C. stylifer Caporiacco, 1935 – Kashmir
C. suruga Nishikawa, 2009 – Japan
C. suthepicus Dankittipakul, Chami-Kranon & Wang, 2005 – Thailand
C. takanawaensis Nishikawa, 2009 – Japan
C. taurus Nishikawa, 2009 – Japan
C. tegenarioides O. Pickard-Cambridge, 1885 – China (Yarkand)
C. tenutubilaris Zhang, Zhu & Wang, 2017 – China
C. terrestris (Wider, 1834) – Europe, Turkey
C. tiantangensis Luo & Chen, 2015 – China
C. tiantongensis Zhang, Peng & Kim, 1997 – China
C. titaniacus Brignoli, 1977 – Greece
C. tochigiensis Nishikawa, 2009 – Japan
C. tojoi Nishikawa, 2009 – Japan
C. tokaraensis Shimojana, 2000 – Japan (Ryukyu Is.)
C. tokunoshimaensis Shimojana, 2000 – Japan (Ryukyu Is.)
C. tominagai Nishikawa, 2009 – Japan
C. tonakiensis Shimojana, 2000 – Japan (Ryukyu Is.)
C. transiliensis Ovtchinnikov, 2000 – Kazakhstan, Kyrgyzstan
C. troglocaecus Shimojana & Nishihira, 2000 – Japan (Okinawa)
C. turkestanicus Ovtchinnikov, 1999 – Russia (Europe) to Central Asia
C. uncatus Liu & Li, 2009 – China
C. undulatus Hu & Wang, 1990 – China
C. unicatus Yaginuma, 1977 – Japan
C. unzenensis Okumura, 2013 – Japan
C. uozumii Nishikawa, 2002 – Japan
C. vallei Brignoli, 1977 – Italy
C. vestigialis Xu & Li, 2007 – China
C. vignai Brignoli, 1978 – Turkey
C. wangi Chen & Zhao, 1997 – China
C. xinjiangensis Hu, 1992 – China
C. yaginumai Nishikawa, 1972 – Japan
C. yahagiensis Nishikawa, 2009 – Japan
C. yambaruensis Shimojana, 2000 – Japan (Ryukyu Is.)
C. yodoensis Nishikawa, 1977 – Japan
C. zaoensis Nishikawa, 2009 – Japan

References

"Gen. Coelotes Blackwall, 1841". World Spider Catalog. Natural History Museum Bern. Retrieved 2019-05-07.
ICZN (1979). "Opinion 1119. Amaurobius C.L. Koch, 1837, and Coelotes Blackwall, 1841 (Araneae): Conserved under the Plenary Powers". Bulletin of Zoological Nomenclature. 35 (4): 216–220.
Lehtinen, P. T. (1967). "Classification of the cribellate spiders and some allied families, with notes on the evolution of the suborder Araneomorpha". Annales Zoologici Fennici. 4: 273.
Blackwall, J. (1841). "The difference in the number of eyes with which spiders are provided proposed as the basis of their distribution into tribes; with descriptions of newly discovered species and the characters of a new family and three new genera of spiders". Transactions of the Linnean Society of London. 18: 601–670. doi:10.1111/j.1095-8339.1838.tb00210.x.