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Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Ordo: Coleoptera
Subordo: Adephaga

Familia: Dytiscidae
Subfamilia: Laccophilinae
Tribus (2): Agabetini - Laccophilini
Overview of genera (13)

Agabetes - Africophilus - Australphilus - Japanolaccophilus - Laccodytes – Laccomimus – Laccophilus - Laccoporus - Laccosternus - Napodytes - Neptosternus - Philaccolilus - Philaccolus - Philodytes


Laccophilinae Gistel, 1856


Alarie, Y.; Nilsson, A.N.; Hendrich, L.; Watts, C.H.S.; Balke, M. 2000: Larval morphology of four genera of Laccophilinae (Coleoptera: Adephaga: Dytiscidae) with an analysis of their phylogenetic relationships. Insect systematics & evolution, 31(2): 121–164. DOI: 10.1163/187631200X00354
Bouchard, P., Bousquet, Y., Davies, A.E., Alonso-Zarazaga, M.A., Lawrence, J.F., Lyal, C.H.C., Newton, A.F., Reid, C.A.M., Schmitt, M., Ślipiński, S.A. & Smith, A.B.T. 2011. Family-group names in Coleoptera (Insecta). ZooKeys 88: 1–972. DOI: 10.3897/zookeys.88.807 Open access. PMC: 3088472 Open access. Reference page.
Braga, R.B. & Ferreira, N. Jr. 2016. Laccomimus xikrin sp. nov. and new records of other species of Laccomimus Toledo & Michat, 2015 (Coleoptera: Dytiscidae: Laccophilinae) from Brazil. Zootaxa 4154(1): 96–100. DOI: 10.11646/zootaxa.4154.1.7. Reference page.
Brancucci, M. 1983: A new genus of the subfamily Laccophilinae (Coleoptera, Dytiscidae). Aquatic insects, 5(4): 251–254. DOI: 10.1080/01650428309361151
Hendrich, L.; Balke, M.; Wewalka, G. 2010: Dytiscidae: Annotated checklist of the Bidessini, Hydrovatini, Hyphydrini, Laccophilinae and Dytiscinae (Coleoptera). Pp. 171-194 in: Jäch, M.A. & Balke, M. (eds.): Water beetles of New Caledonia (part 1). Monographs on Coleoptera, 3. ISSN: 1027-8869 [not seen]
Michat, M.C. 2008: Description of the larvae of three species of Laccophilus Leach and comments on the phylogenetic relationships of the Laccophilinae (Coleoptera: Dytiscidae). Zootaxa, 1922: 47–61. Abstract & excerpt
Toledo, M. & Michat, M.C. 2015. Description of Laccomimus gen. n. and eleven new species from the Neotropical region (Coleoptera, Dytiscidae, Laccophilinae). Zootaxa 3990(3): 301–354. DOI: 10.11646/zootaxa.3990.3.1. Preview (PDF) Reference page.

Laccophilinae is a subfamily of ground and water beetles in the family Dytiscidae. There are at least 410 described species in Laccophilinae.[1][2]


Africophilus Guignot, 1948
Agabetes Crotch, 1873
Australphilus Watts, 1978
Japanolaccophilus Satô, 1972
Laccodytes Régimbart, 1895
Laccophilus Leach, 1815
Laccoporus J. Balfour-Browne, 1939
Laccosternus Brancucci, 1983
Napodytes Steiner, 1981
Neptosternus Sharp, 1882
Philaccolilus Guignot, 1937
Philaccolus Guignot, 1937
Philodytes J. Balfour-Browne, 1939


"Laccophilinae Subfamily Information". Iowa State University. Retrieved 26 January 2018.

"Laccophilinae Report". Integrated Taxonomic Information System. Retrieved 26 January 2018.

D.J. Larson, Y. Alarie, and R.E. Roughley. (2001). Predaceous Diving Beetles (Coleoptera: Dytiscidae) of the Nearctic Region, with emphasis on the fauna of Canada and Alaska. NRC 43253.
Lawrence, J. F., and A. F. Newton Jr. / Pakaluk, James, and Stanislaw Adam Slipinski, eds. (1995). "Families and subfamilies of Coleoptera (with selected genera, notes, references and data on family-group names)". Biology, Phylogeny, and Classification of Coleoptera: Papers Celebrating the 80th Birthday of Roy A. Crowson, vol. 2, 779–1006.
Nilsson, Anders N. (2001). World Catalogue of Insects, volume 3: Dytiscidae (Coleoptera), 395.

Further reading

Arnett, R. H. Jr., and M. C. Thomas. (eds.). (21 December 2000) American Beetles, Volume I: Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia. CRC Press LLC, Boca Raton, Florida. ISBN 978-0-8493-1925-9
Arnett, Ross H. (2000). American Insects: A Handbook of the Insects of America North of Mexico. CRC Press.
Richard E. White. (1983). Peterson Field Guides: Beetles. Houghton Mifflin Company.

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