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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Cladus: Odonatoptera
Cladus: Holodonata
Ordo: Odonata
Subordo: Epiprocta
Infraordo: Anisoptera
Superfamilia: Libelluloidea

Familia: Libellulidae
Subfamilia: Libellulinae
Genus: Libellula
Species: Libellula angelina

Libellula angelina Selys, 1883

IUCN: Libellula angelina Selys, 1883 (Critically Endangered)

Vernacular names
日本語: ベッコウトンボ(鼈甲蜻蛉)
한국어: 대모잠자리

Libellula angelina, also known as bekko tombo, is a species of dragonfly in the family Libellulidae, native to China and Japan. It is under threat by rapid loss of the ponds and other small water bodies it uses as habitat, and is currently classified as critically endangered by the IUCN.


The bekko tombo is a golden to rusty-brown dragonfly, with a dark stripe down the centre of the abdomen and a distinctive pattern of dark markings on its wings. Like other members of the family Libellulidae, this dragonfly has a relatively short, broad abdomen, and its body is distinctly shorter than its wingspan.[2]
Distribution and habitat

The bekko tombo occurs in central and northern China, Japan (Honshu, Shikoku, Kyushu and offshore islands) and Korea.[1] It inhabits old and stable ponds with moderate plant growth in lowland hill areas. It requires clear, open water in which the nymphs crawl around in the bottom sediments.[1][2]

The bekko tombo is univoltine. It has a larval period of one year. Eggs hatch from May to June, and adults emerge the following spring. There is often fierce competition between males for access to reproductive females. Females typically begin to lay eggs in water immediately after copulation, often guarded by their mate. However, females of some species can store live sperm in their body for a number of days.[2]
Threats and Conservation

The bekko tombo has experienced drastic declines in recent decades, mostly driven by the continuing disappearance of its required breeding habitats, old and stable ponds with vegetated borders. Introduced predators such as the largemouth bass (Micropterus salmoides) have also had an impact. The species has been protected in Japan since 1993, and efforts by various NGOs are ongoing to preserve and reconstitute suitable breeding habitats.[2] Current population is estimated at less than 5,000 individuals, and the species has been classified as critically endangered by the IUCN.[1]

Inoue, K. (2006). "Libellula angelina". IUCN Red List of Threatened Species. 2006: e.T11928A3315484. doi:10.2305/IUCN.UK.2006.RLTS.T11928A3315484.en. Retrieved 15 November 2021.
"Bekko tombo (Libellula angelina)". Wildscreen Arkive. Archived from the original on 2015-12-08. Retrieved 5 December 2015.

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