Fine Art

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Cladus: Sarcopterygii
Cladus: Rhipidistia
Cladus: Tetrapodomorpha
Cladus: Eotetrapodiformes
Cladus: Elpistostegalia
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Classis: Reptilia
Cladus: Eureptilia
Cladus: Romeriida
Subclassis: Diapsida
Cladus: Sauria
Infraclassis: Archosauromorpha
Cladus: Crurotarsi
Divisio: Archosauria
Cladus: Avemetatarsalia
Cladus: Ornithodira
Subtaxon: Dinosauromorpha
Cladus: Dinosauriformes
Cladus: Dracohors
Cladus: Dinosauria
Ordo: Saurischia
Cladus: Eusaurischia
Subordo: Theropoda
Cladus: Neotheropoda
Cladus: Averostra
Cladus: Tetanurae
Cladus: Avetheropoda
Cladus: Coelurosauria
Cladus: Tyrannoraptora
Cladus: Maniraptoromorpha
Cladus: Maniraptoriformes
Cladus: Maniraptora
Cladus: Pennaraptora
Cladus: Paraves
Cladus: Eumaniraptora
Cladus: Avialae
Infraclassis: Aves
Cladus: Euavialae
Cladus: Avebrevicauda
Cladus: Pygostylia
Cladus: Ornithothoraces
Cladus: Ornithuromorpha
Cladus: Carinatae
Parvclassis: Neornithes
Cohors: Neognathae
Cladus: Neoaves
Ordo: Bucerotiformes

‎Infraclassis: Aves
Cohors: Neognathae
Cladus: Neoaves
Ordo: Bucerotiformes
Familiae: Bucerotidae – Bucorvidae – Phoeniculidae – Upupidae

Bucerotiformes Fürbringer, 1888: 1567 ["Bucerotes"]
Primary references

Fürbringer, M. 1888. Untersuchungen zur Morphologie und Systematik der Vögel, zugleich ein Beitrag zur Anatomie der Stütz- und Bewegungsorgane. T. van Holkema: Amsterdam. 2 Vols. BHL DOI: 10.5962/bhl.title.51998 Reference page.


Mousebirds, Cuckoo Roller, trogons, hoopoes, hornbills – IOC World Bird List v12.1

Vernacular names
Ελληνικά: Βουκεροτόμορφα
日本語: サイチョウ目
norsk: Hornfugler
polski: Dzioborożcowe

Bucerotiformes /bjuːˈsɛrətɪfɔːrmiːz/ is an order of birds that contains the hornbills, ground hornbills, hoopoes and wood hoopoes.[1] These birds were previously classified as members of Coraciiformes.[2][3][4] The clade is distributed in Africa, Asia, Europe and Melanesia.

Upupidae – hoopoes

Phoeniculidae – wood hoopoes

Bucorvidae – ground hornbills

Bucerotidae – hornbills

Phylogenetic relationships between the families based on a large study by Richard Prum and colleagues published in 2015.[5]

1 Systematics
1.1 Taxonomy
2 References
3 Further reading


Recent genetic data show that ground hornbills and Bycanistes form a clade outside the rest of the hornbill lineage.[6] They are thought to represent an early African lineage, while the rest of Bucerotiformes evolved in Asia. The hoopoe subspecies Saint Helena hoopoe and the Madagascar subspecies are sometimes elevated to a full species. The two wood hoopoe genera, Phoeniculus and Rhinopomastus, appear to have diverged about 10 million years ago, so some systematists treat them as separate subfamilies or even separate families.[7]

Order Bucerotiformes

†Laurillardia smoleni
Suborder Upupi
Family Phoeniculidae
Genus Phoeniculus
Green wood hoopoe, Phoeniculus purpureus
Violet wood hoopoe, Phoeniculus damarensis
Black-billed wood hoopoe, Phoeniculus somaliensis
White-headed wood hoopoe, Phoeniculus bollei
Forest wood hoopoe, Phoeniculus castaneiceps
Genus Rhinopomastus
Black scimitarbill, Rhinopomastus aterrimus
Common scimitarbill, Rhinopomastus cyanomelas
Abyssinian scimitarbill, Rhinopomastus minor
Family Upupidae
Genus Upupa
Eurasian hoopoe, Upupa epops
African hoopoe, Upupa africana
Madagascar hoopoe, Upupa marginata
Suborder Buceroti
Family Bucorvidae
Genus Bucorvus
Abyssinian ground hornbill, Bucorvus abyssinicus
Southern ground hornbill, Bucorvus leadbeateri
Family Bucerotidae
Genus Bycanistes
Trumpeter hornbill, Bycanistes bucinator
Piping hornbill, Bycanistes fistulator
Silvery-cheeked hornbill, Bycanistes brevis
Black-and-white-casqued hornbill, Bycanistes subcylindricus
Brown-cheeked hornbill, Bycanistes cylindricus
White-thighed hornbill, Bycanistes albotibialis
Genus Tropicranus (sometimes included in Tockus)
White-crested hornbill, Tropicranus albocristatus
Genus Tockus
Black dwarf hornbill, Tockus hartlaubi
Red-billed dwarf hornbill, Tockus camurus
Monteiro's hornbill, Tockus monteiri
Red-billed hornbill group
Northern red-billed hornbill, Tockus erythrorhynchus
Damara red-billed hornbill, Tockus damarensis
Southern red-billed hornbill, Tockus rufirostris
Tanzanian red-billed hornbill, Tockus ruahae
Western red-billed hornbill, Tockus kempi
Eastern yellow-billed hornbill, Tockus flavirostris
Southern yellow-billed hornbill, Tockus leucomelas
Jackson's hornbill, Tockus jacksoni
Von der Decken's hornbill, Tockus deckeni
Crowned hornbill, Tockus alboterminatus
Bradfield's hornbill, Tockus bradfieldi
African pied hornbill, Tockus fasciatus
Hemprich's hornbill, Tockus hemprichii
Pale-billed hornbill, Tockus pallidirostris
African grey hornbill, Tockus nasutus
Genus Ocyceros
Malabar grey hornbill, Ocyceros griseus
Sri Lanka grey hornbill, Ocyceros gingalensis
Indian grey hornbill, Ocyceros biostris
Genus Anthracoceros
Malabar pied hornbill, Anthracoceros coronatus
Oriental pied hornbill, Anthracoceros albirostris
Black hornbill, Anthracoceros malayanus
Palawan hornbill, Antracoceros marchei
Sulu hornbill, Anthracoceros montani
Genus Buceros
Rhinoceros hornbill, Buceros rhinoceros
Great hornbill, Buceros bicornis
Rufous hornbill, Buceros hydrocorax
Genus Rhinoplax (sometimes included in Buceros)
Helmeted hornbill, Rhinoplax vigil
Genus Anorrhinus
Austen's brown hornbill, Anorrhinus austeni
Tickell's brown hornbill, Anorrhinus tickelli
Bushy-crested hornbill, Anorrhinus galeritus
Genus Penelopides
Luzon hornbill, Penelopides manillae
Mindoro hornbill, Penelopides mindorensis
Visayan hornbill, Penelopides panini
Samar hornbill, Penelopides samarensis
Mindanao hornbill, Penelopides affinis
Sulawesi hornbill, Penelopides exarhatus
Genus Berenicornis (sometimes included in Aceros)
White-crowned hornbill, Berenicornis comatus
Genus Aceros
Rufous-necked hornbill, Aceros nipalensis
Wrinkled hornbill, Aceros corrugatus
Writhed hornbill, Aceros leucocephalus
Rufous-headed hornbill, Aceros waldeni
Knobbed hornbill, Aceros cassidix
Genus Rhyticeros (sometimes included in Aceros)
Wreathed hornbill, Rhyticeros undulatus
Narcondam hornbill, Rhyticeros narcondami
Sumba hornbill, Rhyticeros everetti
Plain-pouched hornbill, Rhyticeros subruficollis
Papuan hornbill, Rhyticeros plicatus
Genus Ceratogymna
Black-casqued hornbill, Ceratogymna atrata
Yellow-casqued hornbill, Ceratogymna elate


Gill, Frank; Donsker, David, eds. (2019). "Mousebirds, Cuckoo Roller, trogons, hoopoes, hornbills". World Bird List Version 9.2. International Ornithologists' Union. Retrieved 23 July 2019.
Johansson, Ulf S.; Ericson, Per G.P. (2003). "Molecular support for a sister group relationship between Pici and Galbulae (Piciformes sensu Wetmore 1960)" (PDF). J. Avian Biol. 34 (2): 185–197. doi:10.1034/j.1600-048X.2003.03103.x. Retrieved 2008-10-30.
Yuri, T. et al. (2013) Parsimony and Model-Based Analyses of Indels in Avian Nuclear Genes Reveal Congruent and Incongruent Phylogenetic Signals. Biology, 2(1):419-444. doi:10.3390/biology2010419
Jarvis, E.D. et al. (2014) Whole-genome analyses resolve early branches in the tree of life of modern birds. Science, 346(6215):1320-1331.
Prum, R.O.; Berv, J.S.; Dornburg, A.; Field, D.J.; Townsend, J.P.; Lemmon, E.M.; Lemmon, A.R. (2015). "A comprehensive phylogeny of birds (Aves) using targeted next-generation DNA sequencing". Nature. 526 (7574): 569–573. doi:10.1038/nature15697. PMID 26444237.
Woodruff, D. S. & Srikwan, S. 2011. Molecular genetics and the conservation of hornbills in fragmented landscapes. In Poonswad, P. (ed) The Asian Hornbills: Ecology and Conservation. National Center for Genetic Engineering and Biotechnology, Bangkok, pp. 257-264.

Fry, C. Hilary (2003). "Wood-hoopoes". In Perrins, Christopher. The Firefly Encyclopedia of Birds. Firefly Books. p. 383. ISBN 1-55297-777-3.

Further reading

Gonzalez, J.-C.T.; Sheldon, B.C.; Collar, N.J.; Tobias, J.A. (2013). "A comprehensive molecular phylogeny for the hornbills (Aves: Bucerotidae)". Molecular Phylogenetics and Evolution. 67 (2): 468–483. doi:10.1016/j.ympev.2013.02.012. PMID 23438388.

Birds, Fine Art Prints

Birds Images

Biology Encyclopedia

Retrieved from ""
All text is available under the terms of the GNU Free Documentation License

Home - Hellenica World