Circaetus cinereus

Circaetus cinereus (*)

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Cladus: Sarcopterygii
Cladus: Rhipidistia
Cladus: Tetrapodomorpha
Cladus: Eotetrapodiformes
Cladus: Elpistostegalia
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Classis: Reptilia
Cladus: Eureptilia
Cladus: Romeriida
Subclassis: Diapsida
Cladus: Sauria
Infraclassis: Archosauromorpha
Cladus: Crurotarsi
Divisio: Archosauria
Cladus: Avemetatarsalia
Cladus: Ornithodira
Subtaxon: Dinosauromorpha
Cladus: Dinosauriformes
Cladus: Dracohors
Cladus: Dinosauria
Cladus: Saurischia
Cladus: Eusaurischia
Subordo: Theropoda
Cladus: Neotheropoda
Cladus: Averostra
Cladus: Tetanurae
Cladus: Avetheropoda
Cladus: Coelurosauria
Cladus: Tyrannoraptora
Cladus: Maniraptoromorpha
Cladus: Maniraptoriformes
Cladus: Maniraptora
Cladus: Pennaraptora
Cladus: Paraves
Cladus: Eumaniraptora
Cladus: Avialae
Infraclassis: Aves
Cladus: Avebrevicauda
Cladus: Pygostylia
Cladus: Ornithothoraces
Cladus: Ornithuromorpha
Cladus: Carinatae
Parvclassis: Neornithes
Cohors: Neognathae
Cladus: Neoaves
Ordo: Accipitriformes

Familia: Accipitridae
Subfamilia: Circaetinae
Genus: Circaetus
Species: Circaetus cinereus
Name

Circaetus cinereus Vieillot, 1818
References

Vieillot, L.P. 1818. Nouveau Dictionnaire d’Histoire naturelle, appliquée aux arts, à l'agriculture, à l'économie rurale et domestique, à la médecine, etc. Par une société de naturalistes et d'agriculteurs. Avec des figures tirées des trois règnes de la nature. Tome 23. 612 pp. + 6 tt. Déterville, Paris. p. 445 BHL Reference page.

Vernacular names
Afrikaans: Bruinslangarend
čeština: Orlík hnědý
Deutsch: Einfarb-Schlangenadler
English: Brown Snake-Eagle
español: Culebrera Sombría
français: Circaète brun
magyar: Barna kígyászölyv
日本語: チャイロチュウヒワシ
Nederlands: Bruine Slangenarend
polski: Gadożer brunatny
português: Águia-cobreira-castanha
русский: Бурый Змееяд

The brown snake eagle (Circaetus cinereus) is a fairly large species of bird of prey in the family Accipitridae. It is found in West, East and southern Africa. This species is an almost obligate predator of a variety of snakes. A very solitary bird, the brown snake eagle has a prolonged breeding cycle and raises a single eaglet. Although probably naturally scarce, it is classified as a least concern species as it continues to occur over a very broad range.[1]
Distribution and habitat

This species is distributed widely in Africa. In west Africa, ranges from southeastern Mauritania, Senegal, The Gambia, Guinea-Bissau, eastern and western Guinea, Sierra Leone, the northern, inland parts of Liberia, Ivory Coast, Ghana, Togo, Benin, Nigeria as well as inland in Mali and Niger. From there they range across to southern Chad, southern Sudan, South Sudan, inland in Eritrea and northern Ethiopia and central Ethiopia and in southern Somalia through much of Kenya, Uganda and the southern part of the Democratic Republic of the Congo down through southern Africa to about half of Namibia, all of Botswana and Zimbabwe and northeastern and eastern South Africa.[1] Although it is not migratory, brown snake eagles may be somewhat nomadic, with cases of birds on territories stretching up to 200 km (120 mi) apart. Furthermore, ringed birds have been known to travel 2,100 km (1,300 mi) away (from South Africa to the Democratic Republic of the Congo).[2] This species dwell in open woods and wooded savanna, most often preferring areas where gulley or wooded hillocks break up flat areas, apparently preferring somewhat more densely wooded areas than related snake eagles.[3] This species may dwell at any elevation from sea level to 2,000 m (6,600 ft) high.[2]
Description
Adult soaring over Kang in central Botswana
Brown snake eagle hydrating after a drink in Yankari National Park, Bauchi, Nigeria

Their plumage about the body is entirely a fairly dark, hazel brown, with some claims of a purplish sheen in certain light conditions.[2] The body colour extends to the wings but for their contrasting unmarked flight feathers which are whitish-grey. The shortish tail, which is most easily seen in flight, is at all ages barred brown and grayish cream. The juvenile is similar in appearance and colour but tends to have very sparse white feather bases, with birds from south of the range apparently showing heavier white speckling, especially on the abdomen and head.[2][4] This species has a large head and bare legs, which serve to distinguish it from other brownish medium-sized eagles in Africa, although a juvenile bateleur could be confused with one in poor light, but its colour is more varying, its eyes brown and the species has a shorter tail and shorter legs.[2][4] The brown snake eagle is of medium size relative to species referred to as eagles, however it is the largest member of the Circaetus genus. Of the living species in the Circaetinae subfamily, it is similar in size to the bateleur but much smaller than the Philippine eagle. Total length is from 66 to 78 cm (26 to 31 in) and wingspan is from 160 to 185 cm (5 ft 3 in to 6 ft 1 in), though wingspans of up to 200 cm (6 ft 7 in) may possibly be attained.[5][6] Known weights are between 1.5 and 2.5 kg (3.3 and 5.5 lb), with an average of around 2.05 kg (4.5 lb).[7] Despite its fairly large size, this species has a relative small wing spread, being smaller winged than the bateleur and even the much lighter black-breasted snake eagle.[7] The brown snake eagle's call is a hoarse, guttural hok-hok-hok-hok, usually uttered in territorial displays at conspecifics and sometimes culminating in a crowing kaaww. Pairs also call a soft kwee-oo probably as a contact call at the nest.[2]
Biology
Immature in Kruger NP, South Africa.

The brown snake eagle is somewhat larger and more powerful than other snake eagles and consequently tends to take relatively larger prey. It seems virtually any reasonably sized snake, regardless of whether harmless or venomous is taken more or less indiscriminately. Like others in their subfamily, they have natural protection against bites, with thick-skinned legs. [8] 3 out of 4 identified snake species recorded at a nest in Zimbabwe were venomous species, including boomslang, puff adder and black-necked spitting cobra. Most snakes taken here were decapitated before being brought to the nest. Large adult snakes were taken here, including a large puff adder weighing about 750 g (1.65 lb).[9] Even more impressive accounts are known of this species hunting adult black mambas, including specimens measuring up to at least 2.8 m (9 ft 2 in).[4] Hunting is typically from a tree perch or hillock, but also rarely sometimes from flight, and taken almost exclusively on the ground. Like most snake eagles, they quite often swallow their prey whole (though particularly large snakes are torn apart), but seldom do so in flight as do other snake eagles.[2][4][8] Alternate prey is known to have included monitor lizards, toads, francolins, guineafowl and chickens as well as rats and perhaps other mammals.[2][4][9]

This species is solitary and even the breeding pair is rarely seen in the same place at any point of the year. Males do almost all known territorial display flights, which sometimes escalate from typical soaring into butterfly-like erratic flight movements and, if escalated, into interlocking of talons and cartwheeling.[2][4] Breeding takes places in November to July in northern part of range and, although potentially in nearly any month, mainly December-July in Zimbabwe and February-October in Kenya.[2] The nests are relatively small, usually about 60 to 70 cm (24 to 28 in) across and 15 to 30 cm (5.9 to 11.8 in) deep, often being on flat-topped trees such as Acacia or Euphorbia at 3.5 to 11 m (11 to 36 ft) above the ground. Sometimes the nests are infrequently on top of epiphytes, buffalo-weaver nests or on electric pylons.[2][4] Old nests of other raptors are not infrequently used, from tawny eagle to gymnogene nests, and are frequently perceptibly bulkier than those built by snake eagles themselves.[4] A single egg clutch is laid and is incubated mainly by the female for approximately 50 days (quite long for an eagle of this size).[10] Like most birds of prey, the female largely takes on brooding and the male food deliveries. In this species, the male often arrives with a snake hanging from his mouth with only the tail extending from his throat, the female then pulls it out of his bill and throat, thereafter she tears the prey into appropriately sized bites for the single eaglet.[4][8] The juvenile eagles stay around the nest for 60–100 days before exploring branches along the perimeter, until they fledge at 97-113 (mean of about 109) days. The juvenile brown snake eagle is completely independent a few weeks after fledging. Though the young eagle may continue to beg the adults, the parents soon lose interest in feeding the young eagle.[4] Typical lifespan in this species is around 7–10 years, fairly short for an eagle.[4][8] This species is somewhat scarce and is possibly declining overall per the IUCN but it is persisting fairly strongly over a large range, that includes 23.3 thousand square kilometers. It is therefore at least concern status for immediate conservation attention.[1]
References

BirdLife International (2016). "Circaetus cinereus". IUCN Red List of Threatened Species. 2016: e.T22695271A93500155. doi:10.2305/IUCN.UK.2016-3.RLTS.T22695271A93500155.en. Retrieved 12 November 2021.
Ferguson-Lees, J.; Christie, D. (2001). Raptors of the World. Houghton Mifflin Harcourt. ISBN 0-618-12762-3.
Brown, L. H., Fry, C. H., Keithh, S., Urban, E. K., & Newman, K. (1982). The birds of Africa. Academic Press.
Steyn, P. (1983). Birds of prey of southern Africa: Their identification and life histories. Croom Helm, Beckenham (UK). 1983.
del Hoyo, J.; Elliott, A.; Sargatal, J., eds. (1999). Handbook of the Birds of the World. Vol. 2. Barcelona: Lynx Edicions. ISBN 978-84-87334-15-3.
Borrow, N., & Demey, R. (2013). Field guide to the birds of Ghana. Bloomsbury Publishing.
Mendelsohn, J. M., Kemp, A. C., Biggs, H. C., Biggs, R., & Brown, C. J. (1989). Wing areas, wing loadings and wing spans of 66 species of African raptors. Ostrich, 60(1), 35-42.
Steyn, P. (1964). Observations on the Brown Snake-Eagle Circaetus cinereus. Ostrich, 35(1), 22-31.
Irwin, M. P. S. (1981). The birds of Zimbabwe. Salisbury: Quest Publishing.

Tarboton, D. G. (1984). The status and conservation of birds of prey in the Transvaal. Transvaal Museum Monographs, 3(1).

Brown Snake Eagle - Circaetus Cinereus - [1]