Megascops choliba

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Cladus: Sarcopterygii
Cladus: Rhipidistia
Cladus: Tetrapodomorpha
Cladus: Eotetrapodiformes
Cladus: Elpistostegalia
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Classis: Reptilia
Cladus: Eureptilia
Cladus: Romeriida
Subclassis: Diapsida
Cladus: Sauria
Infraclassis: Archosauromorpha
Cladus: Crurotarsi
Divisio: Archosauria
Cladus: Avemetatarsalia
Cladus: Ornithodira
Subtaxon: Dinosauromorpha
Cladus: Dinosauriformes
Cladus: Dracohors
Cladus: Dinosauria
Cladus: Saurischia
Cladus: Eusaurischia
Subordo: Theropoda
Cladus: Neotheropoda
Cladus: Averostra
Cladus: Tetanurae
Cladus: Avetheropoda
Cladus: Coelurosauria
Cladus: Tyrannoraptora
Cladus: Maniraptoromorpha
Cladus: Maniraptoriformes
Cladus: Maniraptora
Cladus: Pennaraptora
Cladus: Paraves
Cladus: Eumaniraptora
Cladus: Avialae
Infraclassis: Aves
Cladus: Avebrevicauda
Cladus: Pygostylia
Cladus: Ornithothoraces
Cladus: Ornithuromorpha
Cladus: Carinatae
Parvclassis: Neornithes
Cohors: Neognathae
Cladus: Neoaves
Ordo: Strigiformes

Familia: Strigidae
Subfamilia: Striginae
Genus: Megascops
Species: Megascops choliba
Subspecies: M. c. choliba – M. c. cruciger – M. c. decussatus – M. c. duidae – M. c. luctisomus – M. c. margaritae – M. c. suturutus – M. c. uruguaiensis – M. c. wetmorei

Synonyms:

M. c. alticola – M. c. caucae – M. c. kelsoi – M. c. guyanensis – M. c. montanus – M. c. portoricensis
[included in cruciger]; M. c. caatingensis
[included in decussatus]; M. c. chapadensis
[included in choliba]; M. c. alilicuco
[included in wetmorei] HBW 2018
Name

Megascops choliba (Vieillot, 1817)
Synonyms

Strix choliba (protonym)
Otus choliba (Vieillot, 1817)

References

Vieillot, L.P. 1817. Nouveau Dictionnaire d’Histoire naturelle, appliquée aux arts, à l'agriculture, à l'économie rurale et domestique, à la médecine, etc. Par une société de naturalistes et d'agriculteurs. Avec des figures tirées des trois règnes de la nature. Tome 7. 586 pp. + 9 tt. Déterville, Paris. p. 39 BHL Reference page.

Vernacular names
English: Tropical Screech-Owl
español: Autillo chóliba
português do Brasil: Corujinha-do-mato

The tropical screech owl (Megascops choliba) is a small species of owl in the family Strigidae. It is found in Costa Rica, Panama, Trinidad, and every mainland South American country.[3]
Taxonomy and systematics

The taxonomy of genus Megascops is somewhat unsettled.[4] A 2015 publication proposed that the tropical screech owl's closest relatives are the white-throated screech owl (M. albogularis) and Koepcke's screech owl (M. koepckeae).[5] However, the International Ornithological Committee (IOC) has not accepted that assessment.[3]

The following nine subspecies are accepted by the IOC, the Clements taxonomy, and BirdLife International's Handbook of the Birds of the World.[3][6][7] M. c. duidae perhaps represents a separate species.[4]

M. c. choliba Vieillot (1817)
M. c. luctisomus Bangs & Pernard (1921)
M. c. margaritae Cory (1915)
M. c. cruciger Spix (1824)
M. c. duidae Chapman (1929)
M. c. decussatus Lichtenstein (1823)
M. c. uruguaii Hekstra (1982)
M. c. surutus Kelso (1941)
M. c. wetmorei Brodkorb (1937)

Description

The tropical screech owl is polymorphic, with much variation in color. It is overall gray-brown or (less commonly) brown or rufous, but always with a relatively distinctive blackish edge to the face and yellow irides. Its undersides are pale with a black "herringbone" pattern. The subspecies differ in the intensity of the colors and the amount of black markings on the upper- and underparts. Adults are 21 to 23 cm (8.3 to 9.1 in) long and weigh 97 to 160 g (3.4 to 5.6 oz).[4]
Distribution and habitat

The tropical screech owl is "the most common and widespread screech-owl of the neotropicis." It is found from Costa Rica and Panama south and throughout South America (including Trinidad and Tobago), almost entirely east of the Andes, but not in the far south. The subspecies are distributed thus:[4]

M. c. choliba, southern Mato Grosso and São Paulo, Brazil, south to eastern Paraguay
M. c. luctisomus, the Pacific slope in Costa Rica south to the Canal Zone in Panama, and on the Pearl Islands
M. c. margaritae, restricted to Margarita Island off of northern Venezuela
M. c. cruciger, Trinidad, and eastern Colombia and Venezuela east to the Guianas and south to eastern Peru
M. c. duidae, Cerro Duida and Cerro de la Neblina in southern Venezuela
M. c. decussatus, central and southern Brazil
M. c. uruguaii, southeastern Brazil, northeastern Argentina, and Uruguay
M. c. surutus, Bolivia
M. c. wetmorei, Paraguay and Argentina

The tropical screech owl inhabits a wide variety of landscapes. Most tend to be partially to mostly open, such as secondary forest, open woodland, coffee plantations, residential areas, and the borders of terra firme and várzea forests. Except for M. c. duidae, it shuns the interior of forests; that subspecies is found in dense forest and bamboo scrub.[4]
Behavior and ecology
Feeding

The tropical screech owl is nocturnal. It forages by sallying from a low perch to capture prey on the ground, from branches, or in the air. Its diet is large arthropods and small vertebrates. The former include worms, scorpions, spiders, and a large variety of insects (such as grasshoppers, moths, cockroaches, cicadas, termites, mantids, crickets, and beetles).[8][9] Vertebrate prey includes frogs, other amphibians, small reptiles[10] (such as snakes),[8] other birds, bats, and rodents.[4]
Breeding

The tropical screech owl's breeding phenology has not been well studied. South of the Equator, the breeding season appears to start in August. The species nests in holes such as natural cavities, old woodpecker holes, and nest boxes. The clutch size is usually one to three but can be as large as six. Adults defend the nest with "fluffing" displays and aggressive moves that may extend to striking humans with their claws.[4]
Vocalization

Dickcissel male perched on a metal pole singing, with neck stretched and beak open.

Songs and calls
Listen to tropical screech owl on xeno-canto

The tropical screech owl's primary song is "short, purring trill, followed by two accentuated clear notes". The female's song is similar to the male's but higher pitched. A secondary, courtship, song is "a bubbling bububúbubu". They also produce "a variety of cackles and longer hooting notes".[4]
Status

The IUCN has assessed the tropical screech owl as being of least concern.[1] It is common in much of its large range, and deforestation may actually benefit it because it provides the more open habitat it frequents.[4]
References
Wikimedia Commons has media related to Megascops choliba.

BirdLife International (2016). "Megascops choliba". IUCN Red List of Threatened Species. 2016: e.T22688774A93208267. doi:10.2305/IUCN.UK.2016-3.RLTS.T22688774A93208267.en. Retrieved 15 August 2021.
"Appendices | CITES". cites.org. Retrieved 2022-01-14.
Gill, F.; Donsker, D.; Rasmussen, P. (July 2021). "IOC World Bird List (v 11.2)". Retrieved July 14, 2021.
Ong, G. (2020). Tropical Screech-Owl (Megascops choliba), version 1.0. In Birds of the World (T. S. Schulenberg, Editor). Cornell Lab of Ornithology, Ithaca, NY, USA. https://doi.org/10.2173/bow.trsowl.01 retrieved August 15, 2021
Remsen, J. V. Jr.; J. I. Areta; E. Bonaccorso; S. Claramunt; A. Jaramillo; D. F. Lane; J. F. Pacheco; M. B. Robbins; F. G. Stiles; K. J. Zimmer (August 2021). "A classification of the bird species of South America". American Ornithological Society. 24. Retrieved August 24, 2021.
Clements, J. F.; T. S. Schulenberg; M. J. Iliff; S. M. Billerman; T. A. Fredericks; B. L. Sullivan; C. L. Wood (2019). "The eBird/Clements Checklist of Birds of the World: v2019". Retrieved August 15, 2019.
"Handbook of the Birds of the World and BirdLife International digital checklist of the birds of the world" (ZIP). HBW and BirdLife International. 5. 2020. Retrieved May 27, 2021.
"Tropical Screech-Owl | the Peregrine Fund".
Proudfoot, Glenn. (2011). Owls of the World, 2nd ed. — Claus König and Friedhelm Weick . 2009. Yale University Press, New Haven, Connecticut.
"Megascops choliba (Tropical Screech Owl)" (PDF). Sta.uwi.edu. Retrieved 10 April 2022.