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Phalacrocorax aristotelis

Phalacrocorax aristotelis (*)

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Cladus: Sarcopterygii
Cladus: Rhipidistia
Cladus: Tetrapodomorpha
Cladus: Eotetrapodiformes
Cladus: Elpistostegalia
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Classis: Reptilia
Cladus: Eureptilia
Cladus: Romeriida
Subclassis: Diapsida
Cladus: Sauria
Infraclassis: Archosauromorpha
Cladus: Crurotarsi
Divisio: Archosauria
Cladus: Avemetatarsalia
Cladus: Ornithodira
Subtaxon: Dinosauromorpha
Cladus: Dinosauriformes
Cladus: Dracohors
Cladus: Dinosauria
Ordo: Saurischia
Cladus: Eusaurischia
Subordo: Theropoda
Cladus: Neotheropoda
Cladus: Averostra
Cladus: Tetanurae
Cladus: Avetheropoda
Cladus: Coelurosauria
Cladus: Tyrannoraptora
Cladus: Maniraptoromorpha
Cladus: Maniraptoriformes
Cladus: Maniraptora
Cladus: Pennaraptora
Cladus: Paraves
Cladus: Eumaniraptora
Cladus: Avialae
Infraclassis: Aves
Cladus: Euavialae
Cladus: Avebrevicauda
Cladus: Pygostylia
Cladus: Ornithothoraces
Cladus: Ornithuromorpha
Cladus: Carinatae
Parvclassis: Neornithes
Cohors: Neognathae
Cladus: Neoaves
Cladus: Aequornithes
Ordo: Suliformes

Familia: Phalacrocoracidae
Genus: Phalacrocorax
Species: P. aristotelis
Subspecies: P. a. aristotelis – P. a. desmarestii – P. a. riggenbachi

Phalacrocorax aristotelis (Linnaeus, 1761)


Pelecanus aristotelis (protonym)
Gulosus aristotelis


Fauna svecica ed.2: XXIII, 51 (#146).

Vernacular names
العربية: غاق
asturianu: Mavea
български: Качулат корморан
brezhoneg: Morvran-guchenn
català: Corb marí emplomallat
corsu: Margonu
čeština: Kormorán chocholatý
Cymraeg: Mulfran Werdd
dansk: Topskarv
Deutsch: Krähenscharbe
Ελληνικά: Θαλασσοκόρακας
English: European Shag
Esperanto: Eŭropa kormorano
español: Cormorán moñudo
eesti: Karikarbas
euskara: Ubarroi mottodun
suomi: Karimetso
føroyskt: Skarvur
français: Cormoran huppé
Gaeilge: Scaga
Gàidhlig: Sgarbh an Sgùmain
galego: Corvo mariño cristado
Gaelg: Arragh Vooar
עברית: קורמורן ימי
hrvatski: Morski Vranac
magyar: Üstökös kárókatona
íslenska: Toppskarfur
italiano: Marangone dal ciuffo
日本語: ヨーロッパヒメウ
ქართული: ქოჩორა ჩვამა
қазақша: Кекілді суқүзғын
lietuvių: Kuoduotasis kormoranas
latviešu: Cekulainais jūras krauklis
македонски: Морски корморан
Malti: Margun tat-Toppu
Nederlands: Kuifaalscholver
norsk nynorsk: Toppskarv
norsk: Toppskarv
polski: Kormoran czubaty
português: Corvo-marinho-de-crista
rumantsch: Cormoran da la cresta
русский: Хохлатый баклан
davvisámegiella: Skárffaš
slovenčina: Kormorán chochlatý
slovenščina: Vranjek
shqip: Karabullaku me çafkë
српски / srpski: Cubasti kormoran - Морски корморан
svenska: Toppskarv
Türkçe: Tepeli karabatak
українська: Чубатий баклан
中文: 歐洲綠鸕鶿

The European shag or common shag (Gulosus aristotelis) is a species of cormorant. It is the only member of the monotypic genus Gulosus.[2] It breeds around the rocky coasts of western and southern Europe, southwest Asia and north Africa, mainly wintering in its breeding range except for the northernmost birds. In Britain this seabird is usually referred to as simply the shag.[3] The scientific genus name derives from the Latin for glutton. The species name aristotelis commemorates the Greek philosopher Aristotle.[4]


The European shag was formerly classified within the genus Phalacrocorax, but a 2014 study found it to be significantly more diverged than the clade containing Phalacrocorax and Urile, but basal to the clade containing Nannopterum and Leucocarbo, and thus classified it in its own genus, Gulosus.[5] The IOC followed this classification in 2021.[6] Gulosus is thought to have split from the Nannopterum-Leucocarbo clade between 9.0 - 11.2 million years ago.[5]

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Young European shag in Croatia
On the nest in Deerness, Orkney

This is a medium-large black bird, 68 to 78 cm (27 to 31 in) long and with a 95-to-110-centimetre (37 to 43 in) wingspan. It has a longish tail and yellow throat-patch. Adults have a small crest in the breeding season. It is distinguished from the great cormorant by its smaller size, lighter build, thinner bill, and, in breeding adults, by the crest and metallic green-tinged sheen on the feathers. Among those differences are that a shag has a lighter, narrower beak; and the juvenile shag has darker underparts. The European shag's tail has 12 feathers, the great cormorant's 14 feathers. The green sheen on the feathers results in the alternative name green cormorant sometimes being given to the European shag.

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Shag in flight

It feeds in the sea, and, unlike the great cormorant, is rare inland. It will winter along any coast that is well-supplied with fish.
Eggs, Collection Museum Wiesbaden

The European shag is one of the deepest divers among the cormorant family. Using depth gauges, European shags have been shown to dive to at least 45 m (148 ft). European shags are preponderantly benthic zone feeders, i.e. they find their prey on the sea bottom. They will eat a wide range of fish but their commonest prey is the sand eel. Shags will travel many kilometres from their roosting sites in order to feed.

In UK coastal waters, dive times are typically around 20 to 45 seconds, with a recovery time of around 15 seconds between dives; this is consistent with aerobic diving, i.e. the bird depends on the oxygen in its lungs and dissolved in its bloodstream during the dive. When they dive, they jump out of the water first to give extra impetus to the dive.

It breeds on coasts, nesting on rocky ledges or in crevices or small caves. The nests are untidy heaps of rotting seaweed or twigs cemented together by the bird's own guano. The nesting season is long, beginning in late February but some nests are not started until May or even later. Three eggs are laid. Their chicks hatch without down and so they rely totally on their parents for warmth, often for a period of two months before they can fly. Fledging may occur at any time from early June to late August, exceptionally to mid-October.

The shag is a pursuit-diving seabird that feeds predominantly in benthic habitats. Due to the relative ease with which diet samples can be collected from this species (regurgitated food or pellets) and the perceived conflict between the Phalacrocoracidae and fisheries, shag diet competition has been the subject of substantial scientific interest.[7][8][9][10] Evidence collected at one colony, the Isle of May, Scotland, between 1985 and 2014, suggests that shag chick diet composition in this population has diversified in response to ocean warming.[11] Shags also feed on fewer sandeel on windy days, presumably due to the strong effect of wind on flight in this species.[12] The year-round diet of full-grown shags at this colony has also changed over the past 3 decades, from sandeel specialists to an increasingly diverse prey base.[13]

There are three subspecies:[14]

G. a. aristotelis – (Linnaeus, 1761): nominate, found in northwestern Europe (Atlantic Ocean coasts)
G. a. desmarestii – (Payraudeau, 1826): found in southern Europe, southwest Asia (Mediterranean and Black Sea coasts)
G. a. riggenbachi – Hartert, 1923: found in northwest African coast

The subspecies differ slightly in bill size and the breast and leg colour of young birds. Recent evidence suggests that birds on the Atlantic coast of southwest Europe are distinct from all three, and may be an as-yet undescribed subspecies.[15]

The name shag is also used in the Southern Hemisphere for several additional species of cormorants.
Example locations
European shags in breeding plumage, Snæfellsnes, Iceland

The European shag can be readily be seen at the following breeding locations in the season (late April to mid July): Saltee Islands, Ireland; Farne Islands, England; Isle of May, Deerness and Fowlsheugh, Scotland; Runde, Norway; Iceland, Denmark; Faroe Islands, Spain; Galicia, Croatia; Dalmatia and Istria. In April 2017, eight new European shags were born in Monaco.[16]

The largest colony of European shags is in the Cíes Islands, Spain, with 2,500 pairs (25% of the world's population).

BirdLife International (2018). "Gulosus aristotelis". IUCN Red List of Threatened Species. 2018: e.T22696894A133538524. doi:10.2305/IUCN.UK.2018-2.RLTS.T22696894A133538524.en. Retrieved 19 November 2021.
Kennedy, Martyn; Spencer, Hamish G. (2014-10-01). "Classification of the cormorants of the world". Molecular Phylogenetics and Evolution. 79: 249–257. doi:10.1016/j.ympev.2014.06.020. ISSN 1055-7903. PMID 24994028.
"Annotated List of the Seabirds of the World - Boobies, Cormorants, Frigatebirds and Tropicbirds". Retrieved 8 October 2007.
Jobling, James A (2010). The Helm Dictionary of Scientific Bird Names. London: Christopher Helm. pp. 55, 301. ISBN 978-1-4081-2501-4.
"Classification of the cormorants of the world". Molecular Phylogenetics and Evolution. 79: 249–257. 2014-10-01. doi:10.1016/j.ympev.2014.06.020. ISSN 1055-7903.
"Taxonomic Updates – IOC World Bird List". Retrieved 2021-07-28.
Snow, Barbara (1960-10-01). "The Breeding Biology of the Shag Phalacrocorax Aristotelis on the Island of Lundy, Bristol Channel". Ibis. 102 (4): 554–575. doi:10.1111/j.1474-919x.1960.tb07132.x. ISSN 1474-919X.
Barrett, RT; Rov, N; Loen, J; Montevecchi, WA (1990). "Diets of shags Phalacrocorax aristotelis and cormorants P carbo in Norway and possible implications for gadoid stock recruitment" (PDF). Marine Ecology Progress Series. 66: 205–218. Bibcode:1990MEPS...66..205B. doi:10.3354/meps066205.
Harris, M. P.; Wanless, S. (1991). "The Importance of the Lesser Sandeel Ammodytes marinus in the Diet of the Shag Phalacrocorax aristotelis". Ornis Scandinavica (Scandinavian Journal of Ornithology). 22 (4): 375–382. doi:10.2307/3676511. JSTOR 3676511.
Cosolo, Mauro; Privileggi, Nicoletta; Cimador, Barbara; Sponza, Stefano (2011-11-01). "Dietary changes of Mediterranean Shags Phalacrocorax aristotelis desmarestii between the breeding and post-breeding seasons in the upper Adriatic Sea". Bird Study. 58 (4): 461–472. doi:10.1080/00063657.2011.603290. ISSN 0006-3657. S2CID 59032510.
Howells, RJ; Burthe, SJ; Green, JA; Harris, MP; Newell, MA; Butler, A; Johns, DG; Carnell, EJ; Wanless, S (2017). "From days to decades: short- and long-term variation in environmental conditions affect offspring diet composition of a marine top predator". Marine Ecology Progress Series. 583: 227–242. Bibcode:2017MEPS..583..227H. doi:10.3354/meps12343.
Lewis, Sue; Phillips, Richard A.; Burthe, Sarah J.; Wanless, Sarah; Daunt, Francis (2015-11-01). "Contrasting responses of male and female foraging effort to year-round wind conditions". Journal of Animal Ecology. 84 (6): 1490–1496. doi:10.1111/1365-2656.12419. ISSN 1365-2656. PMC 4989534. PMID 26283625.
Howells, Richard J.; Burthe, Sarah J.; Green, Jonathan A.; Harris, Michael P.; Newell, Mark A.; Butler, Adam; Wanless, Sarah; Daunt, Francis (2018-11-21). "Pronounced long-term trends in year-round diet composition of the European shag Phalacrocorax aristotelis". Marine Biology. 165 (12). doi:10.1007/s00227-018-3433-9. ISSN 0025-3162.
Gill, Frank; Donsker, David; Rasmussen, Pamela (eds.). "Storks, frigatebirds, boobies, cormorants, darters". IOC World Bird List. Retrieved 30 November 2020.
Yésou, P.; Buchheim, A.; Bermejo, A. (2005). "Which subspecies of Shag occurs in Galicia?" (PDF). British Birds. 98: 369–370.
Verdi, Laurent (April 12, 2017). "Naissance exceptionnelle à Monaco: huit cormorans huppés ont vu le jour". France 3 Côte d'Azur. Retrieved April 13, 2017.

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