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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Cladus: Sarcopterygii
Cladus: Rhipidistia
Cladus: Tetrapodomorpha
Cladus: Eotetrapodiformes
Cladus: Elpistostegalia
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Cladus: Synapsida
Cladus: Eupelycosauria
Cladus: Sphenacodontia
Cladus: Sphenacodontoidea
Cladus: Therapsida
Cladus: Theriodontia
Subordo: Cynodontia
Infraordo: Eucynodontia
Cladus: Probainognathia
Cladus: Prozostrodontia
Cladus: Mammaliaformes
Classis: Mammalia
Subclassis: Trechnotheria
Infraclassis: Zatheria
Supercohors: Theria
Cohors: Eutheria
Infraclassis: Placentalia
Cladus: Boreoeutheria
Superordo: Laurasiatheria
Cladus: †Meridiungulata
Ordo: †Astrapotheria
Familiae: †Astrapotheriidae
Genera: Astraponotus - Astrapothericulus - Astrapotherium - Parastrapotherium - Scaglia - Synastrapotherium - Xenastrapotherium -

Astrapotheriidae is an extinct family of herbivorous South American land mammals that lived from the Late Eocene (Mustersan SALMA) to the Middle Miocene (Laventan SALMA) 37.71 to 15.97 million years ago. The most derived of the astrapotherians, they were also the largest and most specialized mammals in the Tertiary of South America. There are two sister taxa: Eoastrapostylopidae and Trigonostylopidae.

Around 1900, Argentine paleontologist Florentino Ameghino described eight Colhuehuapian (Early Miocene) species from specimens he found south of Lake Colhué Huapi in Patagonia and grouped them into three genera: Parastrapotherium, Astrapotherium, and Astrapothericulus. It was obvious to Ameghino that these species represented a great diversity, ranging in size from a peccary to a rhinoceros, but his description was based entirely on fragmentary and not always comparable dental remains. Other expeditions to Patagonia have subsequently recovered considerably more complete materials.[1]

Genera

According to Kramarz & Bond 2009, Astrapotheriidae includes two clades, Astrapotheriinae and Uruguaytheriinae, and a number of early genera (Astrapotheriidae incertae sedis): Astraponotus (Middle Eocene), Maddenia (Early Oligocene), and Parastrapotherium (Late Oligocene-Early Miocene). Most genera have been found in Patagonia and adjacent areas in Argentina and Chile; whereas members of Uruguaytheriinae have been found further north: Xenastrapotherium (Late Oligocene-Middle Miocene of northern South America), Granastrapotherium (Middle Miocene of Colombia), Uruguaytherium (uncertain age, from Uruguay).[2] According to Kramarz & Bond 2009, the genus Maddenia is a small, pre-Deseadan form of later astrapotheriids.[3] Simpson 1945 grouped Albertogaudrya together with Astraponotus in the subfamily Albertogaudryinae, synonymous with Albertogaudryidae Ameghino 1901.[4] Bond et al. 2011 concluded that a comprehensive evaluation is required regarding astrapotherids.[5]

Incertae sedis
Parastrapotherium Ameghino 1895
Antarctodon Bond et al. 2011
Astrapodon Ameghino 1891
Comahuetherium Kramarz & Bond 2011
Liarthrus Ameghino 1897
Maddenia Kramarz & Bond 2009
Subfamily Albertogaudryinae Simpson 1945
Albertogaudrya Ameghino 1901
Astraponotus Ameghino 1901
Subfamily Astrapotheriinae
Astrapothericulus Ameghino 1901
Astrapotherium Burmeister 1879
Scaglia Simpson 1957
Subfamily Uruguaytheriinae
Uruguaytherium Kraglievich 1928
Granastrapotherium Johnson & Madden 1997
Xenastrapotherium Kraglievich 1928
Hilarcotherium[6]

Notes

Kramaz & Bond 2010, Introduction, p. 183
Goillota et al. 2011, Introduction
Kramarz & Bond 2009, Abstract
Simpson 1945, p. 130
Bond et al. 2011, Relationships

M. C. Vallejo-Pareja; J. D. Carrillo; J. W. Moreno-Bernal; M. Pardo-Jaramillo; D. F. Rodriguez-Gonzalez; J. Muñoz-Duran (2015). "Hilarcotherium castanedaii, gen. et sp. nov., a new Miocene astrapothere (Mammalia, Astrapotheriidae) from the Upper Magdalena Valley, Colombia" (PDF). Journal of Vertebrate Paleontology. 35 (2): e903960. doi:10.1080/02724634.2014.903960. S2CID 130728894.

References

Ameghino, Florentino (1887). Apuntes preliminares sobre algunos mamíferos estinguidos del yacimiento de "Monte Hermoso" existentes en el "Mueso La Plata". Buenos Aires. OCLC 39794328.
Ameghino, Florentino (1891). "Los monos fósiles del Eoceno de la República Argentina" (PDF). Revista Argentina de Historia Natural. Buenos Aires. 1 (6): 383–397. Retrieved 9 March 2013.
Ameghino, Florentino (1895). Première contribution à la connaissance de la faune mammalogique des couches à Pyrotherium. Buenos Aires: P.E. Coni.
Ameghino, Florentino (1897). "Mamiferos Cretaceos de la Argentina. Segunda contribucion al conocimiento de la fauna mastologica de las capas con restos de Pyrotherium". Boletin Instituto Geografico Argentino. 18: 406–521.
Ameghino, Florentino (1901). "Notices préliminaires sur des ongulés des terrains Crétacés de Patagonie". Boletín de la Academia de Ciencias en Córdoba. 16: 349–426. OCLC 123174974.
Bond, Mariano; Kramarz, Alejandro; Macphee, Ross D. E.; Reguero, Marcelo (June 2011). "A New Astrapothere (Mammalia, Meridiungulata) from La Meseta Formation, Seymour (Marambio) Island, and a Reassessment of Previous Records of Antarctic Astrapotheres" (PDF). American Museum Novitates (3718): 1–16. doi:10.1206/3718.2. hdl:2246/6118. OCLC 728156717. S2CID 58908785. Retrieved 9 March 2013.
Burmeister, Hermann (1879). Description physique de la République Argentine : d'après des observations personnelles et étrangères. Vol. 3 Animaux vertébrés, 1. partie, Mammifères vivants et éteints. Paris: Savy. p. 520. OCLC 162707154.
Goillota, Cyrielle; Antoinea, Pierre-Olivier; Tejadab, Julia; Pujosc, François; Gismondid, Rodolfo Salas (2011). "Middle Miocene Uruguaytheriinae (Mammalia, Astrapotheria) from Peruvian Amazonia and a review of the astrapotheriid fossil record in northern South America" (PDF). Geodiversitas. 33 (2): 331–345. doi:10.5252/g2011n2a8. S2CID 129460591. Retrieved 9 March 2013.[permanent dead link]
Johnson, Steven C.; Madden, Richard H. (1997). "Uruguaytheriine Astrapotheres of Tropical South America". In Kay, Richard F.; Madden, Richard H.; Cifelli, Richard L.; Flynn, John J. (eds.). Vertebrate paleontology in the neotropics : the Miocene fauna of La Venta, Colombia. Washington: Smithsonian Institution Press. pp. 355–82. ISBN 9781560984184. OCLC 30320084.
Kraglievich, Lucas (1928). Sobre el supuesto Astrapotherium Christi Stehlin, descubierto en Venezuela (Xenastrapotherium n. gen.) y sus relaciones con Astrapotherium magnum y Uruguaytherium Beaulieui. Buenos Aires: La Editorial Franco-Argentina. OCLC 20881142.
Kramarz, Alejandro G; Bond, Mariano (2009). "A new oligocene astrapothere (Mammalia , Meridiungulata) from Patagonia and a new appraisal of astrapothere phylogeny". Journal of Systematic Palaeontology. 7 (1): 117–128. doi:10.1017/S147720190800268X. S2CID 85351962.
Kramarz, Alejandro; Bond, Mariano (2011). "A new early Miocene astrapotheriid (Mammalia, Astrapotheria) from Northern Patagonia, Argentina". Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen. 260 (3): 277–87. doi:10.1127/0077-7749/2011/0132. OCLC 740850188.
Kramaz, Alejandro G.; Bond, Mariano (2010). "Colhuehuapian Astrapotheriidae (Mammalia) from Gran Barranca south of Lake Colhue-Huapi". In Madden, Richard H.; Carlini, Alfredo A.; Vucetich, Maria Guiomar (eds.). The Paleontology of Gran Barranca: Evolution and Environmental Change Through the Middle Cenozoic of Patagonia. Cambridge University Press. ISBN 9780521872416. Retrieved 9 March 2013.
Simpson, George Gaylord (1945). The principles of classification and a classification of mammals. Bulletin of the AMNH. Vol. 85. New York: American Museum of Natural History. hdl:2246/1104. OCLC 341579.
Simpson, George Gaylord (1957). "A new Casamayoran astrapothere". Revista del Museo Municipal de Ciencias Naturales y Tradicional de Mar del Plata. 1 (3): 11–18. OCLC 81633287.

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