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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Cladus: Sarcopterygii
Cladus: Rhipidistia
Cladus: Tetrapodomorpha
Cladus: Eotetrapodiformes
Cladus: Elpistostegalia
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Cladus: Synapsida
Cladus: Eupelycosauria
Cladus: Sphenacodontia
Cladus: Sphenacodontoidea
Cladus: Therapsida
Cladus: Theriodontia
Subordo: Cynodontia
Infraordo: Eucynodontia
Cladus: Probainognathia
Cladus: Prozostrodontia
Cladus: Mammaliaformes
Classis: Mammalia
Subclassis: Trechnotheria
Infraclassis: Zatheria
Supercohors: Theria
Cohors: Eutheria
Infraclassis: Placentalia
Cladus: Boreoeutheria
Superordo: Euarchontoglires
Ordo: Scandentia

Familia: Tupaiidae
Genus: Dendrogale
Species: Dendrogale melanura
Subspecies (2): D. m. baluensis – D. m. melanura
Name

Dendrogale melanura (Thomas, 1892)

Type locality: Borneo, Sarawak, Mt. Dulit, 5,000 ft.
Vernacular names
References

Dendrogale melanura in Mammal Species of the World.
Wilson, Don E. & Reeder, DeeAnn M. (Editors) 2005. Mammal Species of the World – A Taxonomic and Geographic Reference. Third edition. ISBN 0-8018-8221-4.
Ann. Mag. Nat. Hist., ser. 6, 9: 251.
Wilson, D.E. & Reeder, D.M. (eds.) 2005. Mammal Species of the World: a taxonomic and geographic reference. 3rd edition. The Johns Hopkins University Press: Baltimore. 2 volumes. 2142 pp. ISBN 978-0-8018-8221-0. Reference page. 

The Bornean smooth-tailed treeshrew (Dendrogale melanura) is a species of treeshrew in the family Tupaiidae. It is endemic to Borneo. Its natural habitat is subtropical or tropical dry forests. It is threatened by habitat loss.[2]

Description

Head and body length is 5 inches (13 cm), with tail length 4.5 inches (11 cm). The body mass is about 1.5 oz (43 g). Upper parts of the small body are dark brown, while the under parts are orange-buff with gray bases, and shiny black with reddish streaking along the sides. It has a short snout, with large ear flaps. Prominent orange-brown rings exist around the eyes, with weakly marked facial streaks present on both side of the face, extending from the snout to ears. No shoulder streaks are present. The claws are notably sharp. The thin tail is covered with fine, smooth hair, with darkening towards the tip.[3] D. m. melanura has darker colour above and more reddish below than D. m. baluensis.[4]
Habitat

D. melanura is a terrestrial species. They are diurnal and predominantly arboreal. Live in evergreen rainforest, which active in mossy trees and on rocky boulders in submontane and montane pristine forest. This species seems to feed predominantly on insects.[4]
Distribution

The species is endemic to Borneo, restricted in the mountains of the northwest above 900 m, including the mountain of northeastern Sarawak, Gunung Kinabalu, and Gunung Trus Madi in Sabah. D. m. melanura is recorded from Gunung Dulit, Gunung Mulu, and the Kelabit uplands in northern Sarawak, and from the Sabah-Sarawak border. Meanwhile, D. m. baluensis is recorded from Gunung Kinabalu and Gunung Trus Madi in Sabah.[4]
Status

This species is listed as data deficient because it has not recorded since the early 1970s and it was formerly listed as Vulnerable from 1996 to 2008. The major threats for this species are loss of habitat due to the agricultural expansion and conversion of land to nontree plantations at lower elevations. This species may warrant listing in Near Threatened or higher. The conservation actions only occur at Crocker Range National Park, Sabah, Malaysia.[2]
References

Gardner, A. (2005). Wilson, D.E.; Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Johns Hopkins University Press. pp. 104–105. ISBN 978-0-8018-8221-0. OCLC 62265494.
Gerrie, R.; Kennerley, R. (2019). "Dendrogale melanura". IUCN Red List of Threatened Species. 2019: e.T6405A22278427. doi:10.2305/IUCN.UK.2019-1.RLTS.T6405A22278427.en. Retrieved 26 January 2022.
"Scandentia (Tree Shrews)". Archived from the original on 2009-01-07. Retrieved 2009-02-08.
Francis, C.M. & Payne, J. (2005). A field guide to the mammals of Borneo. Malaysia: Sabah Society

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