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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Cladus: Sarcopterygii
Cladus: Rhipidistia
Cladus: Tetrapodomorpha
Cladus: Eotetrapodiformes
Cladus: Elpistostegalia
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Cladus: Synapsida
Cladus: Eupelycosauria
Cladus: Sphenacodontia
Cladus: Sphenacodontoidea
Cladus: Therapsida
Cladus: Theriodontia
Subordo: Cynodontia
Infraordo: Eucynodontia
Cladus: Probainognathia
Cladus: Prozostrodontia
Cladus: Mammaliaformes
Classis: Mammalia
Subclassis: Trechnotheria
Infraclassis: Zatheria
Supercohors: Theria
Cohors: Eutheria
Infraclassis: Placentalia
Cladus: Boreoeutheria
Superordo: Euarchontoglires
Ordo: Rodentia
Subordo: Myomorpha
Superfamilia: Muroidea

Familia: Cricetidae
Subfamilia: Sigmodontinae
Tribus: Akodontini
Genus: Oxymycterus
Species (16): O. amazonicus – O. caparaoe – O. dasytrichus – O. delatorO. hiskaO. hucuchaO. inca – O. itapeby – O. josei – O. juliacae – O. nasutus – O. nigrifrons – O. paramensis – O. quaestor – O. rufusO. wayku


Oxymycterus Waterhouse, 1837: 21

Type species: Mus nasutus Waterhouse, 1837, by original designation.
Primary references

Waterhouse, G.R. 1837. Characters of new species of the genus Mus, from the collection of Mr. Darwin. Proceedings of the Zoological Society of London 5: 15–21, 27–32.

Additional references

Hershkovitz, P. 1994. The description of a new species of South American hocicudo, or long-nose mouse, genus Oxymycterus (Sigmodontinae, Muroidea), with a critical review of the generic content. Fieldiana: Zoology 79(1460): 1–43. BHL Reference page.
Hoffmann, F.G., Lessa, E.P. & Smith, M.F. 2002. Systematics of Oxymycterus with description of a new species from Uruguay. Journal of Mammalogy 83(2): 408–420. DOI: 10.1644/1545-1542(2002)083<0408:SOOWDO>2.0.CO;2 Online and PDF. Reference page.
Gonçalves, P.R. & Oliveira, J.A. 2004. Morphological and genetic variation between two sympatric forms of Oxymycterus (Rodentia: Sigmodontinae): An evaluation of hypotheses of differentiation within the genus. Journal of Mammalogy 85(1): 148–161. DOI: 10.1644/BER-012
Jayat, J.P., D’Elía, G., Pardiñas, U.F.J., Miotti, M.D. & Ortiz, P.E. 2008. A new species of the genus Oxymycterus (Mammalia: Rodentia: Cricetidae) from the vanishing Yungas of Argentina. Zootaxa 1911: 31–51. Abstract & excerpt Reference page.
Oliveira, J.A. & Gonçalves, P.R. 2015. Genus Oxymycterus Waterhouse, 1837. Pp. 247–268 in Patton, J.L., Pardiñas, U.F.J. & D’Elía, G. (eds.), Mammals of South America. Volume 2. Rodents. The University of Chicago Press: Chicago, Illinois. ISBN 978-0-226-16957-6. Reference page.


Oxymycterus in Mammal Species of the World.
Wilson, Don E. & Reeder, DeeAnn M. (Editors) 2005. Mammal Species of the World – A Taxonomic and Geographic Reference. Third edition. ISBN 0-8018-8221-4.

Vernacular names
English: Hocicudos

Oxymycterus is a genus of rat-like rodents commonly known as hocicudos. They are endemic to South America. As of April 2019, the genus contains the following 16 species:[1]

O. akodontius Thomas, 1921 Argentine hocicudo[2]
O. amazonicus Herzhkovitz, 1994[3] Amazon hocicudo[4]
O. angularis Thomas, 1909 angular hocicudo[5]
O. caparaoe Hershkovitz, 1998 Caparao hocicudo[6]
O. dasytrichus (Schinz, 1821) Atlantic Forest hocicudo[7][8]
O. delator Thomas, 1903 spy hocicudo[9][10]
O. hiska Hinojosa, Anderson & Patton, 1987 small hocicudo[11][12]
O. hispidus Pictet, 1843 hispid hocicudo[12]
O. hucucha Hinojosa, Anderson & Patton, 1987 Quechuan hocicudo[13][12]
O. inca Thomas, 1900 Incan hocicudo[14][15]
O. itapeby Peçanha et al. 2019 Itapevi hocicudo rat[1]
O. josei Hoffmann, Lessa & Smith, 2002[16] Cook's hocicudo[17]
O. nasutus (Waterhouse, 1837) long-nosed hocicudo[18][19]
O. paramensis Paramo hocicudo[20][21]
O. quaestor Thomas, 1903 Quaestor hocicudo[22][23]
O. roberti Thomas, 1901 Robert's hocicudo[23]
O. rufus (Fischer, 1814) red hocicudo[24][25]
O. wayku Jayat et al., 2008 ravine hocicudo[26]


Peçanha, W.T. (April 2019). "A new species of Oxymycterus (Rodentia: Cricetidae: Sigmodontinae) from a transitional area of Cerrado – Atlantic Forest in southeastern Brazil". Journal of Mammalogy. 100 (2): 578–598. doi:10.1093/jmammal/gyz060.
Hershkovitz 1994, pp. 26–27.
Hershkovitz 1994, pp. 23–26.
de Oliveira & Gonçalves 2015, pp. 250–251.
Hershkovitz 1994, pp. 27–26.
de Oliveira & Gonçalves 2015, pp. 251–252.
de Oliveira & Gonçalves 2015, pp. 252–254.
Hershkovitz 1994, p. 28.
de Oliveira & Gonçalves 2015, pp. 254–255.
Hershkovitz 1994, pp. 28–29.
de Oliveira & Gonçalves 2015, pp. 254–256.
Hershkovitz 1994, p. 30.
de Oliveira & Gonçalves 2015, pp. 256–257.
de Oliveira & Gonçalves 2015, pp. 257–258.
Hershkovitz 1994, p. 31.
Hoffmann, Lessa & Smith 2002.
de Oliveira & Gonçalves 2015, pp. 258–259.
de Oliveira & Gonçalves 2015, p. 260.
Hershkovitz 1994, p. 33.
de Oliveira & Gonçalves 2015, pp. 261–263.
Hershkovitz 1994, pp. 33–34.
de Oliveira & Gonçalves 2015, pp. 263–264.
Hershkovitz 1994, p. 34.
de Oliveira & Gonçalves 2015, pp. 264–267.
Hershkovitz 1994, pp. 35–36.

de Oliveira & Gonçalves 2015, pp. 267–268.

Musser, G.G.; Carleton, M.D. (2005). "Superfamily Muroidea". In Wilson, D.E.; Reeder, D.M (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Johns Hopkins University Press. pp. 894–1531. ISBN 978-0-8018-8221-0. OCLC 62265494.
Hershkovitz, Philip (1994). "The Description of a New Species of South American Hocicudo, or Long-Nose Mouth, Genus Oxymycterus (Sigmodontinae, Muroidea), with a Critical Review of Generic Content". Fieldiana: Zoology. New Series. 79: 1–43. doi:10.5962/bhl.title.3351.
Hoffmann, Federico G.; Lessa, Enrique P.; Smith, Margaret F. (2002). "Systematics of Oxymycterus with Description of a New Species from Uruguay". Journal of Mammalogy. 83 (2): 408–420. doi:10.1644/1545-1542(2002)083<0408:SOOWDO>2.0.CO;2.
de Oliveira, João; Gonçalves, Pablo R. (2015). "Gensu Oxymycterus Waterhose, 1837". In Patton, James L.; Pardiñas, Ulyses F. J.; D'Elía, Guillermo (eds.). Rodents. Mammals of South America. Vol. 2. Chicago: The University of Chicago Press. pp. 247–268. doi:10.7208/chicago/9780226169606.001.0001. ISBN 9780226169576.
Jayat, J. Pablo; D'Elía, Guillermo; Pardiñas, Ulyses F. J.; Miotti, M. Daniela; Ortiz, Pablo E. (2008). "A new species of the genus Oxymycterus (Mammalia: Rodentia: Cricetidae) from the vanishing Yungas of Argentina". Zootaxa. 1911 (1): 31–51. doi:10.11646/zootaxa.1911.1.2. ISSN 1175-5334.
Waterhouse, G. B. (1837). "Characteristics of New Species of the Genus Mus, from the Collection of Mr. Darwin". Proceedings of the Zoological Society of London. 5: 15–22. doi:10.1111/j.1469-7998.1837.tb06823.x.

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