Pharsophorus

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Cladus: Sarcopterygii
Cladus: Rhipidistia
Cladus: Tetrapodomorpha
Cladus: Eotetrapodiformes
Cladus: Elpistostegalia
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Cladus: Synapsida
Cladus: Eupelycosauria
Cladus: Sphenacodontia
Cladus: Sphenacodontoidea
Cladus: Therapsida
Cladus: Theriodontia
Cladus: Cynodontia
Cladus: Eucynodontia
Cladus: Probainognathia
Cladus: Prozostrodontia
Cladus: Mammaliaformes
Classis: Mammalia
Subclassis: Trechnotheria
Infraclassis: Zatheria
Supercohors: Theria
Cladus: Metatheria
Ordo: †Sparassodonta

Familia: †Borhyaenidae
Genera: †Pharsophorus
Species: †P. lacrans
Name

Pharsophorus Ameghino, 1897

References
Primary references

Ameghino, A. 1897. Mammiféres crétacés de l’Argentine (Deuxième contribution à la connaissance de la fauna mammalogique de couches à Pyrotherium). Boletin Instituto Geografico Argentino 18(4–9): 406–521.

Pharsophorus is an extinct genus of borhyaenoid sparassodont that inhabited South America during the Middle to Late Oligocene epoch.[1]
Taxonomy

Originally, Pharsophorus was thought to be a borhyaenid, and was even considered to be the ancestor of Borhyaena, Acrocyon, and Arctodictis, but later phylogenetic analyses have shown that it is not a member of the Borhyaenidae and is only more distantly related to these forms.[2][3] Remains of Pharsophorus are known from the Sarmiento Formation of the provinces of Mendoza, Santa Cruz, and Chubut in Argentina, as well as the Salla Formation at the fossil site of Salla in western Bolivia.[4][5] The species "Pharsophorus" antiquus, formerly assigned to this genus, was eventually made the type species of a separate genus Australohyaena.[6]

References

Pharsophorus at Fossilworks.org
Marshall, Larry G. (1978). Evolution of the Borhyaenidae, extinct South American predaceous marsupials. Vol. 117. University of California Press. pp. 1–89. ISBN 9780520095717.
Forasiepi, Analía M. (2009). "Osteology of Arctodictis sinclairi (Mammalia, Metatheria, Sparassodonta) and phylogeny of Cenozoic metatherian carnivores from South America". Monografías del Museo Argentino de Ciencias Naturales. 6: 1–174.
Patterson, Bryan; Larry G. Marshall (1978). "The Deseadan, Early Oligocene, Marsupialia South America". Fieldiana: Geology. 41 (2): 37–100.
Cerdeño, Bryan (2012). "Quebrada Fiera (Mendoza), un importante centro paleobiogeográfico en el Oligoceno tardío de América del Sur". Estudios Geológicos. 67 (2): 375–385. doi:10.3989/egeol.40519.194.
Analía M. Forasiepi, M. Judith Babot and Natalia Zimicz (2014). "Australohyaena antiqua (Mammalia, Metatheria, Sparassodonta), a large predator from the Late Oligocene of Patagonia". Journal of Systematic Palaeontology. 13 (6): 503–525. doi:10.1080/14772019.2014.926403. hdl:11336/59430. S2CID 83669335.