Superregnum: Eukaryota
Cladus: Podiata
Cladus: Amorphea
Cladus: Obazoa
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: ParaHoxozoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Cladus: Olfactores
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Cladus: Sarcopterygii
Cladus: Rhipidistia
Cladus: Tetrapodomorpha
Cladus: Eotetrapodiformes
Cladus: Elpistostegalia
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Cladus: Synapsida
Cladus: Eupelycosauria
Cladus: Metopophora
Cladus: Haptodontiformes
Cladus: Sphenacomorpha
Cladus: Sphenacodontia
Cladus: Therapsida
Cladus: Theriodontia
Cladus: Eutheriodontia
Cladus: Cynodontia
Cladus: Epicynodontia
Cladus: Eucynodontia
Cladus: Probainognathia
Cladus: Prozostrodontia
Cladus: Mammaliamorpha
Cladus: Mammaliaformes
Classis: Mammalia
Cladus: Theriimorpha
Cladus: Theriiformes
Cladus: Trechnotheria
Cladus: Zatheria
Subclassis: Theria
Cladus: Metatheria
Cladus: Marsupialiformes
Infraclassis: Marsupialia
Cladus: Australidelphia
Cladus: Eomarsupialia
Ordo: Diprotodontia
Subordo: Vombatiformes
Familia: Phascolarctidae
Genera (8): Phascolarctos – †Invictokoala – †Litokoala – †Madakoala – †Nimiokoala – †Perikoala – †Priscakoala – ?†Koobor
Name
Phascolarctidae Owen, 1839
Type genus: Phascolarctos Blainville, 1816
References
Additional references
Groves, C.P. 2005. Order Diprotodontia. Pp. 43–70 in Wilson, D.E. & Reeder, D.M. (eds.) . Mammal Species of the World: a taxonomic and geographic reference. 3rd edition. The Johns Hopkins University Press: Baltimore. 2 volumes. ISBN 978-0-8018-8221-0. Reference page.
Archer, M.; Black, K.; Nettle, K. 1997: Giant ringtail possums (Marsupialia, Pseudocheiridae) and giant koalas (Phascolarctidae) from the Late Cainozoic of Australia. Proceedings of the Linnean Society of New South Wales, 117: 3–15.
Black, K. 1999: Diversity and relationships of living and extinct koalas (Phascolarctidae, Marsupialia). Australian mammalogy, 21: 16–17. ISSN: 0310-0049 [not seen]
Black, K.; Archer, M. 1997: Nimiokoala gen. nov. (Marsupialia, Phascolarctidae) from Riversleigh, northwestern Queensland, with a revision of Litokoala. Memoirs of the Queensland Museum, 41: 209–228.
Pledge, N.S. 1992: The Curramulka local fauna: a new Late Tertiary fossil assemblage from Yorke Peninsula, South Australia. Beagle, 9: 115–142. ISSN: 0811-3653
Price, G.J.; Zhao, J-x.; Feng, Y-x.; Hocknull, S.A. 2009: New records of Plio-Pleistocene koalas from Australia: palaeoecological and taxonomic implications. Records of the Australian Museum, 61: 39–48. ISSN: 0067-1975 DOI: 10.3853/j.0067-1975.61.2009.1518
Woodburne, M.O.; Tedford, R.H.; Archer, M.; Pledge, N.S. 1987: Madakoala, a new genus and two species of Miocene koalas (Marsupialia: Phascolarctidae) from South Australia, and a new species of Perikoala. Pp. 293–317 in: Archer, M. (ed.) Possums and opossums: studies in evolution. Surrey Beatty & Sons Pty Limited, England, Chipping Norton. ISBN 0949324051
Vernacular names
español: Fascolárctidos
The Phascolarctidae (φάσκωλος (phaskolos) - pouch or bag, ἄρκτος (arktos) - bear, from the Greek phascolos + arctos meaning pouched bear) is a family of marsupials of the order Diprotodontia, consisting of only one extant species, the koala,[1] and six well-known fossil species, with another six less well known fossil species, and two fossil species of the genus Koobor, whose taxonomy is debatable but are placed in this group. The closest relatives of the Phascolarctidae are the wombats, which comprise the family Vombatidae.[2]
The fossil record of the family dates back to the Middle Miocene[3] or Late Oligocene.[4]
Classification
Family Phascolarctidae[1]
Genus †Nimiokoala
†Nimiokoala greystanesi
Genus †Invictokoala[5]
†Invictokoala monticola
Genus †Madakoala
†Madakoala robustus
†Madakoala wellsi
†Madakoala devisi
Genus †Litokoala
†Litokoala garyjohnstoni
†Litokoala kutjamarpensis
†Litokoala kanunkaensis
Genus †Lumakoala
†Lumakoala blackae[6]
Genus †Koobor
†Koobor jimbarrati
†Koobor notabilis
Genus †Perikoala
†Perikoala palankarinnica
†Perikoala robustus
Genus Phascolarctos
†Phascolarctos maris
Koala - Phascolarctos cinereus
†Giant koala - Phascolarctos stirtoni
†Phascolarctos yorkensis (formerly Cundokoala yorkensis)[7]
Genus †Priscakoala
†Priscakoala lucyturnbullae[4]
References
Groves, C. P. (2005). "Family Phascolarctidae". In Wilson, D. E.; Reeder, D. M (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Johns Hopkins University Press. p. 43. ISBN 978-0-8018-8221-0. OCLC 62265494.
Groves, C. P. (2005). "Suborder Vombatiformes". In Wilson, D. E.; Reeder, D. M (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Johns Hopkins University Press. pp. 43–44. ISBN 978-0-8018-8221-0. OCLC 62265494.
Benton, M.J. (2005). Vertebrate Palaeontology. Oxford. p. 314.
Karen Black; et al. (2012). "New Tertiary koala (Marsupialia, Phascolarctidae) from Riversleigh, Australia, with a revision of phascolarctid phylogenetics, paleoecology, and paleobiodiversity". Journal of Vertebrate Paleontology. 32 (1): 125–138. Bibcode:2012JVPal..32..125B. doi:10.1080/02724634.2012.626825. S2CID 86152273.
Gilbert J. Price and Scott A. Hocknull (2011). "Invictokoala monticola gen. et sp. nov. (Phascolarctidae, Marsupialia), a Pleistocene plesiomorphic koala holdover from Oligocene ancestors". Journal of Systematic Palaeontology. 9 (2): 327–335. Bibcode:2011JSPal...9..327P. doi:10.1080/14772019.2010.504079. S2CID 84316375.
Arthur I. Crichton, Robin M. D. Beck, Aidan M. C. Couzens, Trevor H. Worthy, Aaron B. Camens and Gavin J. Prideaux; et al. (2023). "A probable koala from the Oligocene of central Australia provides insights into early diprotodontian evolution". Scientific Reports. 13 (1). 14521. doi:10.1038/s41598-023-41471-0. PMC 10477348. PMID 37666885.
Talent, John A. (2012). Earth and Life. Springer. p. 1047. ISBN 9789048134281.
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