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Scoteanax rueppellii

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Cladus: Sarcopterygii
Cladus: Rhipidistia
Cladus: Tetrapodomorpha
Cladus: Eotetrapodiformes
Cladus: Elpistostegalia
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Cladus: Synapsida
Cladus: Eupelycosauria
Cladus: Sphenacodontia
Cladus: Sphenacodontoidea
Cladus: Therapsida
Cladus: Theriodontia
Subordo: Cynodontia
Infraordo: Eucynodontia
Cladus: Probainognathia
Cladus: Prozostrodontia
Cladus: Mammaliaformes
Classis: Mammalia
Subclassis: Trechnotheria
Infraclassis: Zatheria
Supercohors: Theria
Cohors: Eutheria
Infraclassis: Placentalia
Cladus: Boreoeutheria
Superordo: Laurasiatheria
Ordo: Chiroptera
Subordo: Microchiroptera
Superfamilia: Vespertilionoidea

Familia: Vespertilionidae
Subfamilia: Vespertilioninae
Tribus: Nycticeiini
Genus: Scoteanax
Species: Scoteanax rueppellii
Name

Scoteanax rueppellii Peters, 1866

Type locality: Australia, New South Wales, Sydney.
Synonyms

australis Iredale (ex MacGillivray), 1937

References

Peters, 1866. Monatsberichte der Koniglich Preussischen Akademie der Wissenschaften zu Berlin, 1866: 21.
Conservation status: IUCN: Scoteanax rueppellii (Near Threatened)
Scoteanax rueppellii in Mammal Species of the World.
Wilson, Don E. & Reeder, DeeAnn M. (Editors) 2005. Mammal Species of the World – A Taxonomic and Geographic Reference. Third edition. ISBN 0-8018-8221-4.

Distribution

East Queensland & East New South Wales (Australia)

Vernacular names
English: Ruppell's Broad-nosed Bat.

Rüppell's or the greater broad-nosed bat (Scoteanax rueppellii) is a species of vespertilionid microbat found in eastern Australia.

Taxonomy

Scoteanax rueppellii was described as a new species in 1866 by German naturalist Wilhelm Peters. The source of the holotype is noted by the author as "Sydney in Westaustralien", referring to Sydney, Australia.[2][4] Peters initially placed it in the genus Nycticejus (alternate spelling of Nycticeius), with the binomial Nycticejus rüppellii. The eponym for the species name "rueppellii" (pronounced rue'-pel-ee-ee) is Eduard Rüppell of the Frankfurt Museum, who loaned the specimen to the author.[2][5] The species was later recognised as sole type of the genus Scoteanax, first described by Ellis Troughton in his standard book of Australian mammals published in 1944.[6][4] The species is distinguished as Rueppell's or the greater 'broad-nosed bat', a term that may refer to species of the genus Scotorepens.[7]
Description

An insectivorous species of bat, moderately large in size with a robust build. The ears of Scoteanax rueppellii are short and widely separated on the head, the tips barely touching if pressed across the head distinguishes physically handled species from similar bats. A concave feature at the outer margin appears below the rounded tip of each ear. The measurements of the forearm is 51 to 56 millimetres, the length of the head and body combined is 63 to 73 mm and tail range is 44–58 mm. The average of the weight range, 21–35 g, is 30 grams. The upper parts of the pelage varies from dark cinnamon to a mid-brown colour, the ventral parts are paler tawny-olive tones. The colour of the patagium forming the wing surfaces is a pinkish shade of brown. The facial features and ears are hairless. A dental features that distinguish S. rueppellii is a single pair of incisors at the upper jaw, immediately adjacent to the canids.[7]

They resemble another species found in eastern Australia, Falsistrellus tasmaniensis (eastern false pipistrelle), which is distinguished by larger ears that overlap by more 5 mm when pressed together, a shorter penis with a large bulb-shaped tip and a gap between the canines and incisors.[7]
Behaviour

Daytime roosts are made within tree hollows and has also been found in urban areas occupying roof structures. The habits and biology of Scoteanax rueppellii are poorly researched and detailed.[8] The species slowly forages at streams or forest edges for larger prey, mainly insects such as flying beetles, or smaller invertebrates, with a languid motion that is restricted in its agility during pursuits.[7][8] The carnivorous diet of this species includes other bats, a behaviour first reported by workers observing the consumption of them in captivity.[9] Scoteanax rueppellii reproduce with a single birth during January.[7]
Distribution and habitat

An uncommon bat which occurs in a wide distribution range along the eastern coast of Australia. At the northern part of the range they occur at the Mount Carbine Tableland, from which the subcoastal range extends south. The preferred habitat is below 500 metres in the southern part of the range, in New South Wales, although the species is only found at higher elevations in the northern tableland regions.[7] The distribution range is poorly understood, and previous assumptions were challenged in a 2006 study that reviewed records and estimated the restrictions to occurrence of the species. The reported constraint by altitude or climatic conditions, or restriction to heavily wooded regions was not supported in the findings, and hypotheses for the distribution for S. rueppellii remain unverified.[10] Scoteanax rueppellii will inhabit wetter gullies of inland forest, but is usually associated with high rainfall regions nearer the coast and with tall forest.[7]
Conservation

As of 2008, it is evaluated as a least-concern species by the IUCN. It meets the criteria for this designation because it has a large geographic range; its population is presumably large; and it is not thought to be experiencing a rapid population decline. However, its population is currently decreasing.[1]
References

Pennay, M. (2020). "Scoteanax rueppellii". IUCN Red List of Threatened Species. 2020: e.T14946A22009127. doi:10.2305/IUCN.UK.2020-2.RLTS.T14946A22009127.en. Retrieved 19 November 2021.
Peters, W. (1866). "Über einige neue oder weniger bekannte Flederthiere". Monatsberichte der Königlichen Preussische Akademie des Wissenschaften zu Berlin. 1866: 16–25.
Troughton, E. le G. 1944. Furred Animals of Australia. Sydney : Angus & Robertson xxviii 374 pp. 25 pls. [353] [publication date determination at Australian Faunal Directory is 1944].
"Species Scoteanax rueppellii (Peters, 1866)". Australian Faunal Directory. Australian Government. 4 December 2018.
Strahan, R.; Conder, P. (2007). Dictionary of Australian and New Guinean Mammals. CSIRO Publishing. p. 139. ISBN 9780643100060.
Jackson, S.M.; Groves, C. (2015). Taxonomy of Australian Mammals. CSIRO Publishing. p. 276. ISBN 9781486300136.
Menkhorst, P.W.; Knight, F. (2011). A field guide to the mammals of Australia (3rd ed.). Melbourne: Oxford University Press. p. ???. ISBN 9780195573954.
"Greater Broad-nosed Bat". The Australian Museum. 12 October 2018.
Churchill, S. (2009). Australian Bats. Allen & Unwin. p. 29. ISBN 9781741766974.
Wilson, P.D. (2006). "The distribution of the greater broad-nose bat Scoteanax rueppellii (Microchiroptera: Vespertilionidae) in relation to climate and topography". Australian Mammalogy. 28 (1): 77–85. doi:10.1071/am06009. ISSN 1836-7402.

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